Tholey Abbey, located in the town of Tholey in Saarland, is widely recognized as Germany’s oldest monastery, with roots tracing back to the 7th century. Over its long history, the abbey has been rebuilt multiple times, with its current Gothic structure dating to the 13th century
The abbey’s most notable recent addition is a set of stained-glass windows designed by the renowned German artist Gerhard Richter in 2020
abstract windows create a striking contrast with the historic surroundings
making Tholey Abbey a unique blend of ancient religious heritage and contemporary art
welcoming both visitors and those seeking a place of worship
These crumbling monastery ruins inspired a master of German Romanticism
This 11th-century Cistercian abbey was once seized by Henry VIII
The remains of one of Wales’ great medieval buildings
The last two abbots of this monastery were both executed by Henry VIII
The remains of a 13th-century Franciscan friary have survived multiple attempts at destruction
The ruins of a 12th-century Cistercian monastery can be found alongside World War II tank traps and pillboxes
An abandoned monastery sits on a far-off island in the Adriatic Sea
image: Gel-based sample pre-fractionation using PEPPI-MS allows for deep proteoform analysis using LC-MS
protein molecules with different chemical structures called proteoforms
are produced from a single gene to perform a variety of physiological protein functions
It has been known for quite some time that humans have approximately 22,000 genes
but the total number of human proteoforms is still unknown
liquid chromatography-mass spectrometry (LC-MS)
a highly sensitive method for measuring biomolecules
is the main method used to analyze proteoforms
Whereas the traditional approach of analyzing proteoforms requires breaking them down into small peptides
a newer approach of analyzing intact proteoforms by LC-MS is called top-down proteomics
Along the lines of the Human Genome Project that mapped all of our genes
there have been attempts to now build an atlas of human proteoforms using top-down proteomics
the proteoform components extracted from biological samples are complex
and LC-MS alone cannot comprehensively detect all proteoforms
the Takemori group at Ehime University originally developed PEPPI-MS
an innovative method for high-resolution fractionation of proteoforms using the inexpensive and simple SDS- PAGE system [Ref
Sample fractionation using PEPPI-MS has been successful in significantly increasing the number of proteoforms detectable by LC-MS
and its performance has led to the adoption of PEPPI in sample preparations
Andreas Tholey’s group at the University of Kiel (Christian-Albrechts-Universität zu Kiel)
an ultra-sensitive proteoform measurement system was built that combines PEPPI-MS fractionation with FAIMS ion mobility mass spectrometry
which can separate proteoforms in the gas phase
they achieved detailed analysis of human cultured cells via top-down proteomics and established a methodology for a new middle-down proteomics
in which the Glu-C digestion products of proteoforms are analyzed [Refs
PEPPI-MS fractionation does not require any special equipment and can be performed with standard biochemical laboratory equipment
Its ease and accessibility has led to its becoming a standard method for sample fractionation in deep top-down proteomics
the Takemori group established experimental protocols for high-resolution proteoform fractionation applying PEPPI-MS and top-down/middle-down proteomics using the FAIMS-LC-MS system and recently published a streamlined protocol combining their work from 2020 to 2024 in Nature Protocols [Ref
The complete set of PEPPI-MS protocols enables high-resolution fractionation of trace biological samples and large-scale proteoform analysis by LC-MS and is expected to contribute to the construction of proteoform atlases for various living species and the development of disease diagnostic methods based on precise proteoform information
10.1038/s41596-024-01100-0
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Volume 11 - 2020 | https://doi.org/10.3389/fpsyg.2020.01885
Lucid dreaming refers to a dream state characterized by the dreamers’ awareness of being in a dream and being able to volitionally control its content
The aim of this study was to describe and model neurophysiological evidence for the seven awareness criteria of lucid dreaming based on those proposed by Paul Tholey
Each of the awareness criteria was analyzed separately with regard to its underlying neurocircuits
but several regions are involved in the state of lucid dreaming
Our results have shown a satisfactory overlap of the awareness criteria and the brain regions activated
that is something other than the sum of its parts
Further research is needed to understand the psychoneurological underpinnings of lucid dreams
Lucid dreaming (LD) is a fascinating research topic and has attracted many enthusiasts
the scientific field is still lacking a comprehensive definition of LD
(3) physical laws need not apply in the dream
(4) the dreamer has a clear memory of the waking world
He declared criteria 1–4 as essential for a “Klartraum,” while criteria 5–7 are optional and do not make a “Klartraum” by themselves
2. Clarity about the freedom of choice (for experiments on the topic see: Libet et al., 1983; Fried et al., 1991; Haggard and Eimer, 1999; Soon et al., 2008; Liljenström, 2015; Liljenström and Nazir, 2016; for an overview see: Baumeister et al., 2010; Caruso, 2012)
The seven criteria used in this article are based on Tholey’s, however, we used an adapted version (Holzinger, 2014) that fits the Gestalt theory terminology better (Yontef, 1993)
We suggest these criteria are more closely related to newer neurophysiological findings and reportings of lucid dreaming experiences
Awareness being a lasting state seems to describe the process of a lucid dream better
compared to a moment of clarity which tends to be momentary
the following criteria are in its core the same as those proposed by Tholey
3. Awareness of (intense) concentration – (awareness of “flow” Csikszentmihalyi et al., 2014)
Awareness of identity (the “I”)
describing this extraordinary state and its potential
The definition of LD is still a work in progress and we hope that the discussion about the definition of a pre-lucid
a lucid dream and Klartraum will gain momentum in the scientific community
and it cannot be reduced to their sum or their difference” (p
It therefore occurs when an entity is observed to have properties its parts do not have on their own and in this case
we assume a model of brain activation on the basis of the seven awareness criteria first described by Paul Tholey
and call it the “lucid brain model,” trying to integrate the varying results of research projects within the last decades
former findings regarding the general difference in brain activity during REM sleep and LD will be discussed
the matter of consciousness in LD will be introduced
and finally neuroscientific evidence for each of our seven proposed awareness criteria will be presented
the conscious experience itself relies on a temporo-parietal-occipital cortical “hot zone.” Therefore
increased activation of the frontal brain regions and temporo-parietal-occipital regions during LD compared to non-LD seem to have numerous effects on conscious awareness
At this point, we would also like to emphasize the notion of consciousness in sleep regarding the understanding and the consequent definition of LD as Harry Hunt did in 1995 (Hunt, 1995) and Jennifer Windt in 2011 (Windt and Noreika, 2011)
We do suspect different stages of consciousness and a lucid dreamer does show higher cognitive abilities and reflective awareness than a non-lucid dreamer overall. Empirical data supports the assumption that LD may be defined as a hybrid state, which is still partially ruled by lower level consciousness (Voss et al., 2009; Dresler et al., 2012; Voss et al., 2018)
This might be the reason that lucid dreams are “happening” as a result of the subconscious
instead of being “created” in the first place
they are a reflection of ourselves and our lives
Both lucid and non-lucid dreams may involve a “thinking” dimension as well as a “perceiving” or “experiencing” dimension
together with the higher activation of meta-cognitive brain areas
possibly supply evidence for the awareness of spatial orientation
This includes the awareness of being in a dream – which is Tholey’s first criteria but is also inherent to our first awareness criteria
The possible activation of all these cognitive processes during LD might explain the awareness of the option to make sound choices based on thoughts
emotions and memories and individual preferences
the concentration of oxygenated hemoglobin (oxy-Hb) was significantly increased in the right and left ventrolateral PFC
They also found a significant increase in oxy-Hb concentration in the right and left dorsolateral PFC
and left ventrolateral PFC while participants were filling out the flow state scale after performing a task in the flow condition
These areas have been found to show increased activation during LD
flow is associated with activity of the PFC
and may therefore be associated with functions such as cognition
the flow experience shares many characteristics with the LD state
We suggest that this area of the brain might also play a role in LD
this is only speculative and requires further exploring
The awareness of the “I” is of course closely related to the awareness of memory
explained in section “Awareness of Memory,” which determines to a great part what the dreamer might decide
wish for or act upon when able to take control of the dream
Together with those findings discussed in section “Awareness of (Spatial) Orientation,” the awareness of the dreaming environment during LD might be explained
General frontal activation might be the reason for the ability to add meaning to a dream by integrating memory
identity and the dreaming environment into a whole
Based on empirical and theoretical findings
we suggest that a dream becomes meaningful by an integration of emotional content (limbic system)
memory (hippothalamus and related structures) and brain structures involved in identity (see section “Awareness of Identity—the “I” Without Which There Would Be No Dialogue”)
This might be possible due to an activation of the DMN and executive functions returning when accessing the state of LD compared to non-LD
together with the anterior cingulate cortex (ACC)
anterior insula and basal ganglia during time perception
as well as time perception might play a role when adding meaning to a dream
Brain regions showing increased activity during lucid REM sleep contrasted with non-lucid REM sleep
Assignment of awareness criteria to brain regions: (1) Awareness of (spatial) orientation: 4
and 8; (2) Awareness of the capacity of choice: 1
and 3; (3) Awareness of (intense) concentration – awareness of “flow”: 1
and 3; (4) Awareness of identity (the “I”) : 4 and 6; (5) Awareness of the dreaming environment : 3
and 8; (6) Awareness of the meaning of the dream : 1
there is still substantial disagreement with regard to the brain regions and frequency bands most activated during lucid dreaming and how they correlate with the theoretical base of lucid dreams
Both authors confirm being the only contributors of this work and approved it for publication
The authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest
We would like to express our sincere thanks to the editor and the reviewers for their careful work and thoughtful suggestions
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Citation: Holzinger B and Mayer L (2020) Lucid Dreaming Brain Network Based on Tholey’s 7 Klartraum Criteria
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†ORCID: Brigitte Holzinger, orcid.org/0000-0001-5385-4091
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Top-down proteomics using mass spectrometry facilitates the identification of intact proteoforms
Multiple past advances have lead to the development of numerous sample preparation workflows
Here we systematically investigated the influence of different sample preparation steps on proteoform and protein identifications
We found that all steps in sample preparation influence the subset of proteoforms identified (for example
physicochemical properties and artificially generated modifications)
The various sample preparation strategies resulted in complementary identifications
substantially increasing the proteome coverage
we identified 13,975 proteoforms from 2,720 proteins of human Caco-2 cells
The results presented can serve as suggestions for designing and adapting top-down proteomics sample preparation strategies to particular research questions
we expect that the sampling bias and modifications identified at the intact protein level will also be useful in improving bottom-up proteomics approaches
the enrichment of smaller proteoforms or the depletion of larger proteoforms is a common step in TDP sample preparation before liquid chromatography with MS/MS (LC–MS/MS) analysis
Upper panel: a suitable LC–MS/MS workflow was established
Middle panel: the established LC–MS/MS workflow was used to investigate the influence of various sample preparation steps on proteoform identification
various prefractionation strategies to enrich proteoforms smaller than 30 kDa and multidimensional separation schemes
Lower panel: data analysis with ProSightPD regarding the proteoform count
physicochemical properties and artificially introduced modifications
two-dimensional reversed-phase low/low pH proteoform fractionation approach
to ensure a fair comparison of different sample preparation strategies
To investigate the influence of various lysis solutions on proteoform and protein identifications by TDP
Caco-2 cells were lysed in (1) phosphate-buffered saline (PBS)
(2) ammonium bicarbonate-buffered urea (Urea–ABC)
(4) Tris-buffered sodium dodecyl sulfate (SDS–Tris)
(5) acidic acetonitrile-water solution containing sodium chloride (ACN–NaCl) and (6) triethylammonium bicarbonate-buffered ACN-water solution (ACN–TEAB)
Number of identified proteoforms (a) and protein accessions (b) (n = 3 replicates of independently performed sample preparations
Full-length proteoforms are those deposited in the proteoform database
including start methionine excision and previously described proteolytic-processed proteoforms
truncated proteoforms refer to all subsequence proteoforms not defined in the database
Overlap coefficients of the proteoforms and proteins identified in the replicates from the various cell lysis conditions
Source data
The lysis with the chaotropic salts urea and GndHCl showed a bias toward more hydrophobic proteins compared to PBS and SDS–Tris lysis
The most similar identifications regarding proteoform and protein overlap coefficient (on average, ~56 and 73%, respectively) were obtained between the Urea–ABC, SDS–Tris and PBS lysis buffers (Fig. 2f)
maintaining a neutral or alkalic pH during the cell lysis
the overlap coefficients with the neutral or alkalic lysis conditions were much lower (45% on proteoform and 67% on protein level)
the lysis with ACN–TEAB and ACN–NaCl resulted in very low overlap coefficients compared to the other lysis conditions (23% on proteoform and 45% on protein level)
Number of identified proteoforms with or without cysteine residues
Fragment maps of selected proteoforms from Glutaredoxin-1 (P35754) identified in the (1) control
(2) reduced and (3) reduced/alkylated samples
The blue brackets represent b-/y-ions identified after CID fragmentation
The cysteine residues highlighted in gray indicate disulfide bridges and those in blue are carbamidomethylated cysteines
Analysis of truncated proteoforms in the control (c) and reduced sample (d)
The two-dimensional histograms display the potential truncation sites of truncated proteoforms
The amino acids N and C terminal of the potential truncation sites are denoted as X and X′
Source data
which potentially reduces the formation of artificially truncated proteoforms
Number of identified full-length and truncated proteoforms (a) and protein accessions (n = 3 replicates of independently performed sample preparations
showing the percentage of proteoforms identified in one
Overlap coefficients between the replicates of all approaches regarding the identified proteins and proteoforms
UpSet plot of proteoforms identified with the various approaches (minimum 50 proteoforms)
Number of identified proteoforms when combining multiple database search results from the different approaches
All raw files were analyzed together in a multiconsensus analysis
and the number of proteoforms identified in a combination of various sample preparations was calculated
C18 material; PEPPI with subsequent MCW precipitation
Source data
only 198 proteoforms (4%) and 163 proteins (12%) were identified in all of the seven investigated approaches
isoelectric point (pI) (b) and GRAVY score (c)
Detection of modifications using MSTopDiff after deconvolution of a randomly selected raw file from each approach with FLASHDeconv
Shown are the intensity × count histograms
Source data
a decent amount of oxidized mass features was detected
which may be attributed to the relatively long sample preparation time
originating from the sample incubation in Laemmli buffer before SDS–PAGE separation
The SPE sample preparation led to the detection of a mass shift assigned to formylation
which is due to the use of formic acid at room temperature
Furthermore, we examined several variations of the investigated prefractionation approaches presented in the Supplementary Results and Supplementary Fig. 17
Both approaches were used to separate proteoforms from human Caco-2 cells in eight fractions, resulting in the identification of a similar number of proteoforms (~2,150) and proteins (~770) (Supplementary Fig. 18a,b)
the two-dimensional separation schemes resulted in more proteoform (~+145%) and protein (~+62%) identifications but required a substantially longer instrument time due to the analysis of eight fractions compared to one sample
providing a comprehensive TDP dataset of human Caco-2 cells
We systematically investigated the influence of different sample preparation procedures on proteoform and protein identifications in TDP
number of replicates and injection amounts
ensured a fair comparison of the different sample preparation steps and can be applied to any TDP study
We showed that each step in the sample preparation influences the number and the physicochemical properties of identifications, that is, biases toward specific proteoform subgroups, such as small, hydrophobic or acidic proteoforms, were introduced depending on the applied approaches (Supplementary Table 7)
we examined the proteoform quality regarding potential artificially introduced modification (for example
due to covalent and noncovalent adducts or peptide bond hydrolysis)
While many sample preparation steps are similar in BUP and TDP
the impact of these steps is often overseen in BUP due to the need for protein inference
we suggest that the results of the intact protein level-centric TDP approach presented here can be used for optimizing BUP workflows
The application of SPE or MWCO allows direct sample cleanup
which may be particularly useful in cases where only limited starting amounts of biological material are available
The findings described here allow us to recommend several guidelines for designing a qualitative TDP study
Three main questions set the frame for the experimental design: (1) what are the objectives of the study
targeting a specific subgroup of proteoforms or reproducible identification of proteoforms with high throughput
(2) How much biological material is available
and what is the time needed for the analysis
(3) How to avoid sample loss and artifacts
Specific examples to explain the use of the decision tree are provided in the Supplementary Information
the selection of the number and combination of parallel approaches depends on the sample amount and instrument time available
should be avoided as this may lead to hydrolysis of peptide bonds C terminal of aspartate residues; (2) the pH value of the cell lysis solution should be neutral or guided by the specific research question
with solution with acidic pH can effectively extract alkaline proteoforms and vice versa; (3) reduction and alkylation of proteoforms should be performed if there is no particular interest in assigning disulfides or other reversible cysteine modifications and (4) possible artificial modifications may be introduced by the cell lysis technique applied
the components of the protease inhibitor or other chemicals used in sample preparation; if possible
reagents that can lead to artificial modifications of proteoforms should be replaced
The data presented can help users make an informed decision on sample preparation based on the specific research objectives
The cOmplete protease inhibitor was from Roche and Acrylamide/Bis solution was from BioRad
Human Caucasian colon adenocarcinoma (Caco-2) cells were from Sigma-Aldrich (cat
fetal bovine serum and TrypLE Express Enzyme were from Thermo Fisher Scientific
all other chemicals were from Sigma-Aldrich
Deionized water (18.2 MΩ cm−1) was prepared by an arium611 VF system (Sartorius)
Human Caco-2 cells were maintained according to the recommendations of the European Collection of Cell Cultures
The cells were cultured at 37 °C and 5% CO2 in RPMI-1640 medium (25 mM HEPES
0.013 mM phenol red) supplemented with 10% (v/v) fetal bovine serum
When a confluence between 90 and 100% was reached
the cells were passaged by detaching with TrypLE Express Enzyme
the cells were washed three times with PBS (200g
The cells were stored at −70 °C until cell lysis and further use
Caco-2 cells were lysed in 1× PBS (pH 7.4) supplemented with 1× cOmplete protease inhibitor by ultrasonication (5 × 20 s on ice)
the protein concentration was determined by Pierce BCA assays (Thermo Fisher
Scientific) following the manufacturer’s instructions
protein determination was performed by BCA assay (Thermo Fisher
and proteoforms were purified by MCW precipitation
and injected twice using the LMW and HMW methods
For protein purification, MCW precipitation was used9
Cell lysates containing 100 or 500 µg of protein were made up to 125 µl with MilliQ and 400 µl of methanol
100 µl of chloroform and 275 µl of MilliQ were added
The samples were vortexed and centrifuged for 10 min at 14,000g
The upper phase was removed and 600 µl of methanol was added
and the protein pellet was washed twice with 600 µl of methanol
The protein pellet was dried in a fume hood and stored at −20 °C until further processing
To investigate the influence of cysteine reduction or alkylation on proteoform identifications
Caco-2 cells were lysed and the proteoforms were aliquoted and subjected in triplicates to reduction with or without subsequent alkylation
Caco-2 cells were lysed in 8 M GndHCl and 200 mM triethylammonium bicarbonate (pH 8.5) supplemented with 1× cOmplete protease inhibitor
The proteoforms (100 µg in 50 µl of lysis buffer) were reduced by adding 1.6 µl of 200 mM tris(2-carboxyethyl)phosphine and incubation at 50 °C for 1 h
alkylation was performed by adding 3.3 µl of 375 mM iodoacetamide and incubating for 30 min at room temperature in the dark
the proteoforms were purified by MCW precipitation
100 µg of Caco-2 proteins were purified by MCW precipitation
The proteoforms were resuspended in 200 µl of LC–MS loading buffer (3% ACN
Note that a considerable proportion of the pellet could not be dissolved and was visible as an insoluble precipitate
the samples were centrifuged for 20 min at 4 °C
For the enrichment of proteoforms below approximately 30 or 50 kDa
appropriate MWCO filters were used (Amicon Ultra-0.5
500 µg of Caco-2 proteins were precipitated by MCW precipitation and resuspended in 500 µl of 8 M GndHCl
The MWCO filters were equilibrated by adding 500 µl of 8 M GndHCl and centrifugation for 15 min at room temperature
The filters were then transferred to a new reaction vial
The filters were centrifuged for approximately 15 min at 14,000g until 50 µl of the samples remained on the filter
The filter was centrifuged again after adding 8 M GndHCl to a total volume of 500 µl
was purified with an equilibrated 3 kDa MWCO filter
the sample was centrifuged at 14,000g for 30 min
and the flowthrough was discarded (residue in the filter was approximately 50 µl)
the sample was washed twice with 100 mM TEAB (pH 8.5) on the filter
The residual containing the re-buffered proteoforms was transferred to a new 1.5-ml reaction tube
where the filter was inverted and centrifuged at 2,000g for 3 min
The sample was lyophilized and solubilized in 50 µl of LC–MS loading buffer
one band below approximately 30 kDa of two adjacent lanes was excised based on the prestained protein marker without previous fixation or staining of the gel
After incubation for 10 min at 20 °C and 1,500 rpm
the gel pieces were removed using a CoStar-X filter (2,000g
the solutions were subjected to 3-kDa MWCO filters and washed (13,500g
25 min) twice with 8 M urea and 100 mM ABC
the proteoforms were purified by anion-exchange material via the stage-tip protocol
The stage tip was conditioned with 40 µl of methanol and equilibrated with 120 µl of 100 mM ABC (7,000g
The sample was loaded on the stage tip and washed with 120 µl of 100 mM ABC
The proteoforms were eluted with 80 µl of 50% ethanol and 0.5% formic acid
The sample was lyophilized and solubilized in 20 µl of LC–MS loading buffer
The proteoform fractionation by GELFrEE (Expedeon) was performed according to the manufacturer’s recommendations
500 µg of Caco-2 proteins were purified by MCW precipitation and resuspended in 30 µl GELFrEE loading buffer
112 µl of MilliQ and 8 µl of 1 M dithiothreitol
The sample was incubated for 10 min at 50 °C before loading in the sample loading chamber
For the enrichment of proteoforms smaller than approximately 30 kDa
an 8% Tris-Acetate cartridge was used and the separation was performed according to the manufacturer’s protocol
Only the first fraction containing proteoforms smaller than ~30 kDa was purified by MCW precipitation and resuspended in 50 µl of LC–MS loading buffer before LC–MS/MS analysis
The sample was lyophilized and resuspended in 20 µl of LC–MS loading buffer before LC–MS/MS analysis
500 µg of Caco-2 proteins were purified by MCW precipitation and resuspended in 150 µl 8 M GndHCl
the sample was centrifuged for 20 min at 21,100g
and the supernatant was transferred into a new reaction vial
50 mg; PerkinElmer) cartridge was used for SPE
The SPE material was activated by 2× 1 ml 100% ACN and equilibrated by 2× 1 ml 5% formic acid before sample loading
The sample was washed twice with 5% formic acid and eluted with 300 µl of 70% ACN and 300 µl of 100% ACN
The proteins were resuspended in 50 µl of LC–MS loading buffer before LC–MS/MS analysis
500 µg of Caco-2 proteins were purified by MCW precipitation
and 76% ACN supplemented with 50 mM sodium chloride
The small protein fraction was resuspended by vigorous vortexing
ultrasonication and incubation for 1 h at 20 °C and 1,500 rpm
20 °C) to remove the insoluble (large) proteoforms
and the supernatant was transferred in a new reaction vial
100 µl of MilliQ was added and the sample was frozen at −80 °C
The proteins were resuspended in 20 µl of LC–MS loading buffer before being subjected to LC–MS/MS analysis
The fractions were vacuum-dried by lyophilization and resuspended in 40 µl of LC–MS loading buffer
The pools were dried by lyophilization and resuspended in 20 µl of LC–MS loading buffer before LC–MS/MS analysis
The processing steps were combined in a multiconsensus search for merging multiple datasets
The C-score is a metric that defines well-characterized proteoforms and stringent filtering allows the minimization of false-positive proteoform identifications
In the Supplementary Notes
various observations that notoriously occur during sample preparation and LC–MS/MS analysis are presented
which can provide valuable insights into potential issues to consider when designing TDP experiments
The GRAVY score is a measure for the hydrophobicity of the proteoforms: that is
The term annotated proteoforms refers to full-length proteoforms deposited in the database
and previously described truncated proteoforms
subsequence proteoforms are all truncated ones not defined in the database
The potential truncation sites were determined by matching the truncated proteoform sequence to the full-length sequence deposited in the database
Further information on research design is available in the Nature Portfolio Reporting Summary linked to this article
Proteoform: a single term describing protein complexity
Proteomes are of proteoforms: embracing the complexity
Seeing the complete picture: proteins in top-down mass spectrometry
Deep top-down proteomics revealed significant proteoform-level differences between metastatic and nonmetastatic colorectal cancer cells
The Blood Proteoform Atlas: a reference map of proteoforms in human hematopoietic cells
Mapping the proteoform landscape of five human tissues
Laser capture microdissection-capillary zone electrophoresis-tandem mass spectrometry (LCM-CZE-MS/MS) for spatially resolved top-down proteomics: a pilot study of zebrafish brain
Quantitative top-down proteomics in complex samples using protein-level tandem mass tag labeling
Best practices and benchmarks for intact protein analysis for top-down mass spectrometry
On-capillary cell lysis enables top-down proteomic analysis of single mammalian cells by CE-MS/MS
Novel strategies to address the challenges in top-down proteomics
Comparing top-down proteoform identification: deconvolution
Multidimensional separations in top–down proteomics
On the scalability and requirements of whole protein mass spectrometry
Top-down proteomics and the challenges of true proteoform characterization
Characterization of large intact protein ions by mass spectrometry: what directions should we follow
Enhancing top-down proteomics of brain tissue with FAIMS
Mass spectrometry-based top-down and bottom-up approaches for proteomic analysis of the Moroccan Buthus occitanus scorpion venom
PEPPI-MS: polyacrylamide-gel-based prefractionation for analysis of intact proteoforms and protein complexes by mass spectrometry
Recent advances in top-down proteome sample processing ahead of MS analysis
Optimization of a top-down proteomics platform for closely related pathogenic bacterial discrimination
Coupling high-field asymmetric waveform ion mobility spectrometry with capillary zone electrophoresis-tandem mass spectrometry for top-down proteomics
Sensitive top-down proteomics analysis of a low number of mammalian cells using a nanodroplet sample processing platform
Multiplexed size separation of intact proteins in solution phase for mass spectrometry
Solid phase extraction as sample pretreatment method for top-down quantitative analysis of low molecular weight proteins from biological samples using liquid chromatography—triple quadrupole mass spectrometry
Multidimensional separation schemes enhance the identification and molecular characterization of low molecular weight proteomes and short open reading frame-encoded peptides in top-down proteomics
A top-down proteomics platform coupling serial size exclusion chromatography and Fourier transform ion cyclotron resonance mass spectrometry
Deep top-down proteomics using capillary zone electrophoresis-tandem mass spectrometry: identification of 5700 proteoforms from the Escherichia coli proteome
Identification of proteoforms by top-down proteomics using two-dimensional low/low pH reversed-phase liquid chromatography-mass spectrometry
Development of an online 2D ultrahigh-pressure nano-LC system for high-pH and low-pH reversed phase separation in top-down proteomics
Two-dimensional separation using high-pH and low-pH reversed phase liquid chromatography for top-down proteomics
Improved identification of proteoforms in top-down proteomics using FAIMS with internal CV stepping
Improved label-free quantification of intact proteoforms using field asymmetric ion mobility spectrometry
Top-down ion mobility/mass spectrometry reveals enzyme specificity: separation and sequencing of isomeric proteoforms
Spatially resolved top-down proteomics of tissue sections based on a microfluidic nanodroplet sample preparation platform
The C-score: a Bayesian framework to sharply improve proteoform scoring in high-throughput top down proteomics
Sense and nonsense of elevated column temperature in proteomic bottom-up LC-MS analyses
Validation of top-down proteomics data by bottom-up-based N-terminomics reveals pitfalls in top-down-based terminomics workflows
Nonproteolytic cleavage of aspartyl proline bonds in the cellulosomal scaffoldin subunit from Clostridium thermocellum
Depletion of high-molecular-mass proteins for the identification of small proteins and short open reading frame encoded peptides in cellular proteomes
high-quality feature deconvolution for top-down proteomics
MSTopDiff: a tool for the visualization of mass shifts in deconvoluted top-down proteomics data for the database-independent detection of protein modifications
Artifactual isoform profile modification following treatment of human plasma or serum with protease inhibitor
monitored by 2-dimensional electrophoresis and mass spectrometry
Site-specific pyrolysis-induced cleavage at aspartic acid residue in peptides and proteins
Optimization of protein-level tandem mass tag (TMT) labeling conditions in complex samples with top-down proteomics
Toward a universal sample preparation method for denaturing top-down proteomics of complex proteomes
Accurate estimation of context-dependent false discovery rates in top-down proteomics
Size-based proteome fractionation through polyacrylamide gel electrophoresis combined with LC–FAIMS–MS for in-depth top-down proteomics
Orbitrap mass spectrometry and high-field asymmetric waveform ion mobility spectrometry (FAIMS) enable the in-depth analysis of human serum proteoforms
Revealing Corynebacterium glutamicum proteoforms through top-down proteomics
Advancing top-down analysis of the human proteome using a benchtop quadrupole-orbitrap mass spectrometer
UniProt: the Universal Protein Knowledgebase in 2023
PyOpenMS: a Python-based interface to the OpenMS mass-spectrometry algorithm library
Pyteomics 4.0: five years of development of a Python proteomics framework
ProteomeXchange provides globally coordinated proteomics data submission and dissemination
Download references
Steinbach for cultivating the human Caco-2 cells
This work was supported by the DFG Cluster of Excellence ‘Precision Medicine in Inflammation’
decision to publish or preparation of the manuscript
Open access funding provided by Christian-Albrechts-Universität zu Kiel
Systematic Proteome Research and Bioanalytics
Institute for Bioinformatics and Medical Informatics
performed the experiments and LC–MS/MS measurements
wrote the original draft of the manuscript
review and editing of the final manuscript
The authors declare no competing interests
Nature Methods thanks Claire Boos, Alexander Leitner and the other, anonymous, reviewer(s) for their contribution to the peer review of this work. Peer reviewer reports are available
in collaboration with the Nature Methods team
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Supplementary Table 5 Proteoforms identified
Supplementary Table 6 Proteins with associates proteoforms identified
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The small village of Tholey in southwestern Germany is home to the oldest monastery on German soil. Founded in the seventh century, the Abbey of St Maurice was plundered and badly damaged 1000 years later during the wars of the French Revolution, and only re-established in 1949. But local philanthropists have breathed new life into the old walls.
The highlight of the extensive works are the stained glass windows for the north and south aisles of the abbey church by Afghanistan-born artist Mahbuba Maqsoodi
and the three choir windows designed by Germany’s most famous living artist
The art establishment are calling this the most significant project of its kind on the continent of Europe
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activities on preceding journals at which the article was previously under consideration are not shown (for instance submission
All content on this site: Copyright © 2025 Elsevier B.V.
This article contains an account of an attempted suicide
If you or a loved one is thinking about suicide
call or text the National Suicide Prevention Lifeline at 988
Crystal Tetu had a deep relationship with her grandmother
"I was raised by my grandparents," said Tetu
Very structured — we always had dinner together
As Tholey aged and moved into an independent living facility
Tetu said Tholey was known for showing new residents around and helping them feel comfortable
"She just took it upon herself to help out new people."
She was hospitalized several times for heart failure
"This happened in May and June and the end of July," Tetu said
and I missed that — she didn't tell me because she didn't want to ruin my honeymoon."
Tholey ultimately told Tetu that she wasn't improving
A problem arose: A doctor explained to Tetu that Tholey was no longer getting nutrients from food
Her body was no longer digesting food because it needed that energy to keep her heart beating
"She's not going to get better," Tetu said
"and she was wondering why she had to suffer like that
She just had to stick it out until she starved to death."
because I strongly believe that my grandma shouldn't have had to suffer that way," Tetu said
"There's suffering that is required as part of being alive
Tetu said she supports medical aid in dying — also known as physician-assisted suicide and assisted death — a practice that is legal in 10 U.S
a bill is in play at the Minnesota Legislature this year
The potential passage of a MAID law in Minnesota has inspired passionate testimony from those who support — and those who oppose — the practice
"My grandma should have had the option," Tetu said
"It's modeled after the Oregon Death with Dignity law that's been in effect since 1997," said Dr
a Minneapolis physician and Compassion & Choices' Doctors for Dignity program director
"They must be able to self-ingest the medication through the GI tract
Medical aid in dying is currently legal in 10 states and the District of Columbia
Oregon was the first state to enact MAID policy in 1997
have based their MAID laws off of Oregon's
the national campaign strategist for policy reform organization Death with Dignity
are among the co-authors for Minnesota's End-of-Life Option Act in their respective chambers
said he first heard about medical aid in dying from a constituent on the campaign trail in 2022
just through discussions about this being something that was important to them," Smith said
how tragic some of these end-of-life moments are
this great law in place that allowed some of these more horrific situations to be alleviated."
she said her support for the bill comes through the lens of bodily autonomy
have experienced patients' deaths," Boldon said
"and I know what that can look like and how awful that can be for some people
for people wanting to make their own decisions about what that looks like for them."
has introduced MAID legislation several times since 2016
the End-of-Life Option Act has gained more traction compared to previous years
is the longest bill hearing I've had so far as I've been in the Legislature
and tragic hearing about people's loss of close family members and very difficult
and it has since passed the House's public safety and judiciary committees and will be considered by the Commerce Finance and Policy Committee next
"We've had it through the health committee
which is a big barrier because the health committee is very limited on time," Smith said
says a lot about what she thinks about the issue and what can be done with that bill this session."
Boldon said she would be surprised if the Senate's End-of-Life Option Act (SF 1813) received a committee hearing due to time constraints and not having enough support to pass
"It's something that I very much hope we can continue talking about because it's important
but there is a diversity of thought around this within my caucus in the Senate," Boldon said
"And so I don't see it having the votes to pass this session."
Thoman said whether this bill makes it to Gov
Tim Walz's desk depends on "whether Minnesotans get out there and start talking about how important this is to them."
"Lawmakers have said they're dedicated to respect for bodily autonomy
privacy in medical decision-makings," Thoman continued
"And so this fits right along with their priorities and their values."
Not all Minnesotans who have publicly commented on the End-of-Life Option Act have been in favor of the proposal
there were three testifiers who said they live in Rochester
the healthy and the sick," said Karin Charron of Rochester
"The catechism of the Catholic Church says we are stewards
Many testifiers — from a group of 40 Muslim doctors to the National Association of Catholic Nurses — shared Charron's opposition to MAID on religious and moral grounds
"From our standpoint (at Death with Dignity)," Sugerman said
"forcing all people in the country to adhere to a particular religious belief is just not the way we should be operating."
at least three Rochester physicians submitted written testimony for the MAID bill
"I believe this would be a significant detriment to the way medicine is practiced in Minnesota," wrote Dr
"The supporters of this bill are undoubtedly well-intentioned in their desire to relieve suffering for terminally ill patients
the means that they would legalize are highly unethical and should not be condoned by the state."
That impact on practicing medicine is something the Minnesota Alliance for Ethical Healthcare is concerned about — in a statement
alliance president Nancy Utoft said the proposal "introduces risks and unintended consequences for Minnesotans who already face severe health care disparities and marginalization
people of color and people with limited income."
"This proposed legislation would turn the practice of medicine on its head by creating a new standard of care that views assisted suicide the same as treatments that ease pain and increase quality of life," Utoft continued
"It would force providers to inform patients of their option to die or to refer patients to a provider that will prescribe the lethal dosage
How are patients — especially those who have previously experienced marginalization in health care — supposed to trust that doctors are always working in their best interests if this is a requirement of our care system?"
Physicians and other medical providers are able to opt out of providing MAID to a patient who requests it
some doctors have voiced concern over having to present MAID as an option to patients even if they would not offer that option
"The only requirement is that if a patient requests that their medical records be transferred
that that's done in a timely manner," Thoman said
"That's really the only requirement for health professionals or organizations in this bill."
No matter how Minnesota's End-of-Life Option Act shakes out this year
the issue of medical aid in dying is something that will stay on Julianne Donaldson's mind
until 2015 when they moved to Dundas to be closer to their hobby farm
Don was diagnosed with esophageal and stomach cancer
in which she asked her husband if he — faced with a dire diagnosis — ever thought of committing suicide
I would never do something like that,'" Donaldson said
we had a grandson who took his own life ..
and until you experience something like that
I don't think anyone can realize the depth of that action
Don went through chemotherapy and radiation
it became apparent that "he was going to lose his life to that horrible cancer."
Donaldson said she was brushing her teeth when she heard Don
I'm going to shoot myself,'" Donaldson said
you're not,' and I turned him around and got him back in bed."
After a conversation with a social worker and a nurse
she learned what had changed for Don: the physical and mental pain was overwhelming
"I didn't know it would be this hard," she remembers him saying
and the hospice caretakers did so for the next two weeks until he passed away on Jan
Donaldson now advocates for MAID legalization in Minnesota
they should be allowed that bodily autonomy
especially when there is no hope," Donaldson said
it would have been so much of an easier passing for him and for us
The years leading up to 1861 saw polarised politics
One of the US’s foremost historians reflects on America’s Disunion – then and now
“I tremble for my country when I reflect that God is just,” Thomas Jefferson wrote in 1781. The American revolution still raged, many of his own slaves had escaped, his beloved Virginia teetered on social and political chaos
who had crafted the Declaration of Independence for this fledgling nation at war with the world’s strongest empire
felt deeply worried about whether his new country could survive with slavery
“daily exercised in tyranny,” with slaveholders practicing “unremitting despotism”
He admitted that slaveholding rendered his own class depraved “despots” and destroyed the “amor patriae” of their bondsmen
“Can the liberties of a nation be thought secure when we have removed their only firm basis
a conviction in the minds of the people that these liberties are of the gift of God?” This advocate of the natural rights tradition
ended his rumination with the vague entreaty that his countrymen “be contented to hope” that a “mollifying” of the conditions of slaves and a new “spirit” from the revolution would in the “order of events” save his country
Danger cannot come from abroad … If destruction be our lot
we must ourselves be its author and finisherAbraham Lincoln in 1838For that republic to survive it took far more than hope and a faith in progress
it did not survive; in roughly four score years it tore itself asunder over the issue of racial slavery
as well as over fateful contradictions in its constitution
and the reimagining of the second republic that resulted form the pivot of American history
The civil war sits like the giant sleeping dragon of American history ever ready to rise up when we do not expect it and strike us with unbearable fire
It has happened here – existential civil war
fought with unspeakable death and suffering for fundamentally different visions of the future
as our first and second founders well understood
Americans love to believe their history is blessed and exceptional
the story of a people with creeds born of the Enlightenment that will govern the worst of human nature and inspire our “better angels” to hold us together
But this most diverse nation in the world is still an experiment
and we are once again in a political condition that has made us ask if we are on the verge of some kind of new civil conflict
Abraham Lincoln worried about politicians’ unbridled ambition
and about the “perpetuation of our political institutions”
The abolitionist Elijah Lovejoy had just been murdered by a mob the previous year in Illinois
Lincoln saw an “ill omen” across the land due to the slavery question
He felt a deep sense of responsibility inherited from the “fathers” of the revolution
How to preserve and renew “the edifice of liberty and equal rights”
“At what point shall we expect the approach of danger?” Lincoln asked
“By what means shall we fortify against it?” His worries made him turn inward
“Shall we expect some transatlantic military giant to step the ocean
Asia and Africa combined … could not by force
in a trial of a thousand years.” Lincoln did not fear foreign enemies
we must ourselves be its author and finisher
Those words were prescient in Lincoln’s own century
But they have a frightful clarity even today
Are Americans on the verge of some kind of social disintegration
as the writer Paul Starobin has recently asked
in which he revisits the old thesis that the secession moment represented a “crisis of fear” that led tragically to disunion and war
It remains to be seen whether we have a deep enough well of tolerance and faith in free speech to endure this “catharsis” we seem to seek
View image in fullscreenFar-right protesters clash with anti-fascist demonstrators in Charlottesville
Photograph: Michael N/Pacific/BarcroftImagesPsychological explanations
do not fully explain America’s current political condition
We are in conflict about real and divergent ideas
Are we engaged in a “cold civil war” as one writer has suggested
with so much rhetorical and real violence in the air
have many serious people using words like “unprecedented”
or phrases like “where in time are we” or “we haven’t been here before”
Commentators and ordinary citizens have been asking how or where in the past we can find parallels for our current condition
Trump has been the gift that keeps on giving
his flouting of political and constitutional traditions
his own party’s moral bankruptcy in its inability to confront him
have forced the media to endlessly ask historians for help
That moral cowardice by Republicans shows some glimmers of hope; Mitt Romney has just called out Trump
accusing him of “unraveling … our national fabric” by his coziness with white supremacists
and Senator Bob Corker of Tennessee charged Trump with putting the nation “in great peril” by his incompetence and racism
Sixteen years ago, in the book, Race and Reunion: The Civil War in American Memory
I made a simple claim: “As long as America has a politics of race
it will have a politics of civil war memory.” Unfortunately
as well as conferences and courses taught on the same subject
that prescription seems truer now than ever
The line from the killings of Travon Martin and Michael Brown
through a myriad of other police shootings
and then especially from the mass murder of nine African Americans in Charleston in June
to the recent white supremacist demonstration and violence in Charlottesville mark a dizzying
America is in the midst of yet another of its racial reckonings which always confront us with a shock of events we are
Just now we are engaged in a frenzied wave of Confederate monument removals; it is a manifestation of how well-meaning Americans can demonstrate their anti-racism and full of admirable impulses
But this too in all likelihood will not itself prepare us for the next shock of events nor our next reckoning
All parallels are unsteady or untrustworthy
But the present is always embedded in the past
the fateful decade that led to the civil war
After the Great Hunger in Ireland the US experienced an unprecedented immigration wave between 1845 and the mid-1850s
prompting a rapid and powerful rise of nativism
Irish and German Catholics were unwelcome and worse
the nation’s first expansionist foreign conflict
stimulated an explosive political struggle over the expansion of slavery
The Fugitive Slave Act of 1850 caused a wave of “refugee” former slaves escaping the northern states into Canada
as well as a widespread crisis over violent rescues of fugitive slaves
the constant flight of slaves from the South to free states was
became increasingly politicized as it became more radical
Americans had to ask whether their institutions would last
either disintegrated or broke into sectional parts
Third parties suddenly emerged with success like no other time in our history
whose xenophobia and anti-Catholicism got them elected in droves in New England in the early 1850s
And the most successful third party in our history
were born in direct resistance to the Kansas-Nebraska Act of 1854
and which opened up the western territories to the perpetual expansion of slavery
A succession of weak and pro-slavery presidents from 1844 through 1860 either tarnished the institution of the presidency or deepened the sectional and partisan divide
the supreme court weighed in by declaring in Dred Scott v Sandford that blacks were not and could never be citizens of the US
“for more than a century been regarded as beings of an inferior order … so far inferior that they had no rights which the white man was bound to respect.” This most notorious court decision legally opened up all of the west
all of the north to the presence of slavery
So discredited was the supreme court among many northerners in the wake of the decision that the Republicans made resistance to the judiciary a rallying cry of their political insurgency
That impulse led to the election of Lincoln in 1860
who firmly controlled that region’s politics
as the primary impulse to secede from the union
They believed they could not co-exist in a nation now led by a political organization devoted to their destruction
By the time of the sectionalized and polarized election of 1860
conducted in a climate of violence and danger caused by John Brown’s raid on Harper’s Ferry in 1859
north and south had developed broad-based mutual conspiracy theories of each other
They did so through a thriving and highly partisan press
Both sides tended to have their own sets of facts and their own conceptions of both history and the constitution
White southerners feared and loathed abolitionists
and now they faced anti-slavery politicians who could truly affect power and legislation if elected
pro-slavery interests had developed a widespread theory about a “black Republican” conspiracy in the north
determined on taking hold of all reins of government to put slavery
one southern leader after another pronounced against what they perceived as an abolitionist conspiracy against their livelihoods and their lives
the secession commissioner for Mississippi
1860 that Republicans “now demand equality between the white and negro races
under our constitution; equality in representation
equality in the right of suffrage … equality in the social circle
equality in the rights of matrimony.” He concluded therefore
the deep south faced a stark choice: “Sink or swim
she will never submit to the principles and policy of this black Republican administration.”
advanced an equally potent idea of a “slave power” conspiracy that had grown into a staple of antislavery politics
consisted of the southern slaveholding political class; they were obsessively bent on control of every level of government and every institution – presidency
The slave power especially demanded control over future expansion of the United States in order for its system to survive
The theory made greater sense with time to many people
was increasingly a minority interest in the federal government
No one made this case about the slave power better than the black abolitionist Frederick Douglass
1853 Douglass gave the slave power clear definition
It was “a purely slavery party” in national affairs and its branches reached “far and wide in church and state”
The conspiracy’s “cardinal objects” were suppression of abolitionist speech
removal of free blacks from the United States
the “nationalization” of slavery in every state of the union
and the expansion of slavery to Mexico and South America
Douglass saw the slave power as an all-encompassing national plague with “instinctive rapacity”
with a “natural craving after human flesh and blood”
It was a “murderous onslaught” upon the rights of all Americans to sustain the claims of a few
would be “thawing a deadly viper instead of killing it”
He had faith in the “monster’s” inherent tendency to over-reach and destroy itself
“it is also crushing itself.” It had “made such a frightful noise” with the “Fugitive Slave Act… the Nebraska bill
the recent marauding movements of the oligarchy in Kansas,” that it now performed as the abolitionists’ “most potent ally”
Douglass detected a great change in northern public opinion
Instead of regarding the abolitionists as mere fanatics “crying wolf”
the masses now perceived the evil in their midst and themselves cried “kill the wolf”
Thus we might see one of the strongest parallels of all between the road to disunion and our current predicament
The rhetoric about the slave power and about black Republicans has a familiar ring today
Millions of Americans on the right who garner their information from selective websites
radio shows and Fox News possess all sorts conspiratorial conceptions of liberals and the alleged radical views of professors on university campuses
Many on the left also know precious little about people in rural and suburban America who voted for Trump; coastal elites do sometimes hold contemptuous views bordering on the conspiratorial about the people they “fly over”
Americans are more than politically polarized; we are bitterly divided about our expanding diversity
about the right to vote and how to protect it
over whether we even believe in a social contract between citizens and the polity
we are divided over conflicting visions of our future
Let us hope that we find ways to fight out our current conflicts within politics and not between each other in our over-armed society
the white supremacist far right will become its own worst enemy
As Americans consider the survival of their own amor patriae we might reflect on just how old our story is
When Moses sent the Israelites across the Jordan
he instructed them to put up memory stones to mark their journey and their story
Americans have put up more than their share of memory stones
and are just now living through a profound process of deciding which ones will remain
But as we look deeply into just what our own amor patriae means
we might think hard about what inscriptions we want written on the memory stones of our own times
We might draw one from Douglass in 1867: “We ought to have our government so shaped that even when in the hands of a bad man we shall be safe.”
View upcoming auction estimates and receive personalized email alerts for the artists you follow
Gerhard Richter
Gerhard Richter has completed his final major artwork
On the occasion of the unveiling of three stained-glass windows in the Tholey Abbey in the Saarland in South Germany
which were produced at the direction of the world’s foremost painter
he spoke to the Deutsche Presse-Agentur (German press agency) and said the windows were surely his last ‘work number.’ Starting with Tisch in 1962
all of Richter’s works are registered in a directory
which will now be complete with the windows assigned the number 957
The new Gerhard Richter stained glass windows in the Tholey Abbey
The good news is that this does not mean that his creation will cease altogether
While his age of 88 doesn’t permit Richter to continue with painting large-scale artworks
he will create a new directory for drawings and sketches that he is now dedicating himself to
“Not so great,” the coy and humble painter told German press
The windows aren’t the first of their kind by Richter
the “Richter-Fenster,’ created by use of 72 colors and a random pattern of small squares
The execution of the Tholey Abbey windows was carried out by the Gustva van Treeck glass workshop in Munich
Both the window patterns stand in close relation to paintings Richter has made in the past
his 4096 Colours in the case of the Cologne window
and motives from his book Patterns in the newly unveiled one
It is not by accident that Richter came to design church windows with their unique effects
Many of his abstract paintings in their colorful harmony evoke this spiritual effect
like the meeting of a setting sun with the colorful stained glass of a house of prayer
The majority of Richter’s work is dedicated to abstraction
but also to photorealistic paintings and his own kind of surrealism
His oeuvre stands somewhere between Modernist Art and developed into the Contemporary
but he stands as living witness to a period that has largely faded
The secondary market has long endowed his masterpieces with eight-figure values
albeit the figures for such a major persona in the chronicles of art history still come in short compared to some of his contemporaries and successors
but the test of time might change that balance
# of Lots Offered and Sold: Gerhard Richter
The most expensive recorded artwork by the German
sold for $46.4 million at Sotheby’s in 2015
establishing a new record for an artwork by a living artist
earning Richter’s dismay at the astronomical sums spent on his art
The record was broken the same year still by Jeff Koons’ Balloon Dog (Orange)
but Richter’s paintings are still the fourth
fifth and sixth most expensive works by a living artist
The mania in the first half of the previous decade surrounding his works has cooled slightly in the last years
evident by the decline of yearly performances in the second half
This is not to say that an average yearly sales value north of $150 million is not substantial
but the records of 2012 and 2014 with $293.8 million and $289.6 million
Not that he would have any ambition to do so
the master painter was granted another special honor
when Sotheby’s named its summer concluding sale Rembrandt to Richter
demonstrating his importance in a sale filled with major names of 500 years of art history
The monumentally-sized Wolken (Fenster) only reservedly fulfilled its expectations
as it achieved $13.5 million after a $11.6 million - $15.5 million estimate
This year’s high is expected to be broken on October 06
when Sotheby’s will offer Abstraktes Bild (649-2) at its Hong Kong Contemporary Sale with an estimate of $15.3 million - $17.9 million
is not looking to break records and is and was largely interested in art for its own sake
It is surely a large part of why he has achieved such depth and brilliance and is revered by the world
The completion of his last major work is therefore a sad development for the art world
but a fitting and well-deserved one for an artist who has been creating for almost 60 years.
For more on auctions, exhibitions, and current trends, visit our Magazine Page
By Flora Graham
(Image: EURAC Institute for Mummies and the Iceman)
When peering at these magnified blood cells, you’re not just looking closer than the naked eye can – you’re also looking through the mists of time. The dark areas on the cells show bruising that occurred over 5000 years ago, when Ötzi the iceman met his end on the Schnalstal glacier in the Alps
Ötzi seems to have died just hours after sustaining serious injuries in hand-to-hand combat. Now Frank Maixner from the Institute for Mummies and the Iceman in Bolzano
and Andreas Tholey from Kiel University in Germany have found evidence that he received a blow to the forehead that caused his brain to knock against the back of his skull
the blood cell samples were extracted through tiny holes that already existed
By investigating the cells’ proteins
the researchers were able to see a snapshot of Ötzi’s last hours
regardless of where it originates in the body
whereas proteins provide precise information about what is happening in specific regions within the body,” says Tholey
“When you think that we have succeeded in identifying actual tissue changes in a human who lived over 5000 years ago
you can begin to understand how pleased we are.”
Ötzi has already been photographed more than the most paparazzi-loved celebrity – the Iceman photoscan website puts many of these images online
Journal reference: Cellular and Molecular Life Sciences, DOI: 10.1007/s00018-013-1360-y
Clarification: When this article was first published
the Leipzig-based startup behind the vegan chocolate and protein brands Nucao and Numove
has just bagged €14 million in a Series A round of financing
Proceeds from the round will fuel Nu’s ongoing expansion
amid a record-breaking year of sales for the company
driven by rising consumer demand for health-conscious and plant-based foods.
German startup The Nu Company
has announced the closing of its Series A funding with more than €14 million
also saw participation from Paris-headquartered food tech investors Five Seasons Ventures and DX Ventures
the VC arm of food delivery giant Delivery Hero
Other notable investors include former racing driver and Formula 1 world racing champion Nico Rosberg
who has since his retirement backed a number of green tech startups like Tier Mobility
as well as BackBone Ventures and the Austrian F&B investors Square One Foods
which is the startup’s first financing round
will help it grow its distribution across Europe and internationally.
“This Series A will help us bring more products to market that can bring about a real and positive change in the food industry,” said co-founder and CMO Christian Fenner
“We want our products to be even easier to access throughout Europe
whether at the local supermarket or with a single click online.”
Fenner launched the startup back in 2016 with Thomas Stoffels and Mathias Tholey
and it has since rolled out two successful consumer-facing brands
plastic-free and low-sugar vegan chocolate and protein bars and snacks
and free from refined sugars and packaged in home compostable cellulose-based wrappers.
For every product sold, Nu sets aside a portion of proceeds to support reforestation projects led by nonprofit organisation Eden
Nu’s funding round comes amid surging demand for more sustainable and healthy foods, especially climate-friendly snacks, which is quickly becoming a big trend in the food industry
Nu itself recorded triple-digit sales growth for 2021 for the fourth consecutive year
and the brand is now retailed in more than 10,000 stores
Co-founder and CFO Stoffels says that the startup is now on track to plant more than 10 million trees before the end of 2021.
“We don‘t stop at our own products. There needs to be a change in consciousness in the industry as a whole,” said Tholey, who issued with his co-founders an open call for climate action on the part of German politicians and corporations
“We’re putting the pressure on and demanding that politicians and large corporations follow suit
so we can treat consumers and our planet with respect and operate in a way that’s appropriate for all generations.”
Nu’s purpose-driven business is a major part of why it believes the brand is poised for growth amid changing consumer trends
“The goal is always to bring a healthier option to everyone’s table without neglecting the environment,” said principal Clément Helinckx.
“The result is excellent consumer engagement and outstanding growth rates,” Helinckx continued
“It’s exciting to see a brand spread its wings so quickly
This shows a clear adjustment of the market and a great deal of potential to trigger long-term changes.”
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