Metrics details
Tiger sharks are apex predators with a circumglobal tropical and warm-temperate distribution
with a general lack of population data for the central Indian Ocean
tiger sharks display frequent use of the harbour area
and reproductive characteristics of the world’s largest known tiger shark aggregation in a geographically-restricted area
photo identification and laser photogrammetry
we identified 239 individual tiger sharks over a 7-year study period
The aggregation was female-dominated (84.5%)
with both large juveniles and adults present
Adult females were resighted over the entire study period displaying strong inter- and intra-annual site fidelity
Modelled residency using maximum likelihood methods suggests they spent 60.7 ± S.E
shorter residence periods and longer absence periods compared to juvenile females
Prolonged abdominal distensions of adult females indicate they likely stay near Fuvahmulah during gestation and reproduce biennially
Fuvahmulah seems to provide suitable conditions for gestation given the year-round provision of food and warm waters
exhibited by strong site fidelity and temporal residency
Our results show indications of a thriving population within the confines of protected waters
Location of Fuvahmulah within the Maldives Archipelago in the Indian Ocean. The dive sites, where tiger sharks are frequently spotted and where most footage originates from, are marked in the map (Map was created using QGIS 3.28.1-Firenze, URL: https://qgis.org/download/)
With a lack of data throughout the rest of the Indo-Pacific and the possibility of localised populations
the reproductive cycle length remains unclear
particularly for tiger sharks in the Maldives
this study represents the first extensive investigation of the species with this method
this study presents the first scientific description of this tiger shark aggregation
with implications for the management and conservation of tiger sharks in the Maldives
Between Dec 7th 2016 and Sep 30th 2023, we collected footage from a total of 788 separate dive surveys: 772 at the Tiger Harbor (TH) dive site, six at Farikede (FK) and ten from Offshore Plateau (OP) (Fig. 1)
We saved 32,495 photographs and frame grabs from video material of sufficient quality to identify individual sharks
(a) Countershading delineation in six different tiger sharks
F-025) display intraspecific variation of the countershading delineation anterior to the pectoral fins
This feature was most useful in differentiating the individuals
(b) Cumulative number of identifications of tiger sharks by sex per month visiting tiger harbour
The orange line indicates the cumulative number of dive surveys per month
visual size estimates were assumed to be sufficient for approximate TL estimation of the tiger sharks
Total length (TL) of visual size estimates from 213 sharks incremented by 0.5 m
All females 3.5 m and larger were considered adults
(a) Standardised width of presumably pregnant sharks over time
Day 0 indicates the last measurement before the sharks’ absence period
where we presume parturition may take place
Sharks scored 1 or 0 were visually assessed as ‘pregnant’ or ‘not pregnant’
while ‘not scored’ refers to a shark’s appearance
where we did not infer pregnancies based on their visual appearance (see Methods for more details)
(b) An example of one individual’s presumed pregnancy (F-049)
Number in brackets provides a corresponding day value to a)
The standardised width of this shark increased until day 0 followed by a period of absence for 93d
the standardised width of this shark had significantly declined
Two consecutive pregnancies were recorded for nine individuals and three consecutive pregnancies were recorded for three individuals
Consecutive periods of gestation and parturition were not significantly different from an expected two-year period (Chi-squared test
Footage of pregnant sharks was separated on average by 788.7 (SD = 69.0) days or 2.16 years
The residency models were run for (i) the entire population, (ii) adult females, and (iii) juvenile females. Model H, which included parameters for emigration, reimmigration and mortality of tiger sharks, had the lowest QAIC indicating the best fit for the data of all three data sets (Table 1)
overall tiger sharks spent a mean of 65.5 ± S.E
76.1 [95%CI (55.5–79.4)] days in Fuvahmulah and a mean of 107.9 ± S.E
Aggregation size was estimated to be 43.1 ± S.E
3.3 [95%CI (37.2–50.4)] individuals present in the study area on any given day
adult female tiger sharks spent a mean of 60.7 ± S.E
7.5 [95% CI (50.2–72.9)] days in Fuvahmulah and a mean of 110.4 ± S.E
Aggregation size was estimated to be 25.9 ± S.E
2.2 [95% CI (22.0–30.5)] individuals present in the study area on any given day
juvenile female tiger sharks spent a mean of 93.0 ± S.E
42.7 [95% CI (32.7–182.1)] days in Fuvahmulah and a mean of 76.9 ± S.E
whereas the aggregation size was estimated to be 16.0 ± S.E
2.1 [95% CI (12.0–20.2)] individuals present in the study area on any given day
juvenile females spent shorter time periods away from the study site than adults
while remaining for longer periods when present
Their overall aggregation size was also considerably lower
which is consistent with the amount of juveniles vs
The results for all tiger sharks was similar to the results for the adult females except for aggregation size
mean ± S.E.) for all tiger sharks (top panel) and for juvenile (red) and adult (blue) female tiger sharks visiting Fuvahmulah
Model H (pale lines) is represented in all cases including emigration
reimmigration and mortality as model presets
The LIR plot for juvenile females showed high probability of resighting from day 1 to 6 days. LIR remained high until 687 days, or 1.9 years. After 2,158 days the LIR reached zero, implying permanent emigration or mortality (Fig. 5)
most juveniles have not been identified for periods longer than three years
The LIR plot for adult females displayed a rapid decline from day 1 to 189 days
Afterwards the model curve stabilised at a value of 0.014
the LIR increased significantly and roughly maintained this level until 1457 days
indicating strong periodicity in the visitation of the dive site by some adult female tiger sharks as well as inter-annual site fidelity
This study provides the first assessment of the population structure
reproductive patterns and residency behaviour of tiger sharks at a hotspot in the central Indian Ocean using non-invasive methods
While adult females showed inter- and intra-annual site fidelity with temporal residency periods
large juveniles showed high residency with shorter periods of absence suggesting that they possibly remain resident in close geographical proximity to the island
Fuvahmulah hosts an unprecedentedly large aggregation of tiger sharks year-round
which appears to play a critical role for the population’s reproductive cycle i.e
This low-effort food source probably attracted tiger sharks to the island long before the tourism activities started and altered their distribution in the area
we assume that this large aggregation is a consequence of human activity and that sharks within this study likely present a skewed picture of natural tiger shark population dynamics
we recognize that there exists a level of ambiguity regarding the classifications made in this study
Our adoption of > 3.0 m TL as a length-at-maturity is a conservative estimate
more likely to categorise adult individuals ≤ 3.0 m as juveniles rather than the converse
Tunas with massive chunks missing are occasionally observed in the local fishing market
likely due to depredation events by tiger sharks (Supplementary Fig
these events have not been quantified for the tuna fishery in Fuvahmulah
we recommend avoiding generalisations of reproductive cycle lengths within the Indo-Pacific
the occurrence of tiger sharks at TH is biased by intraspecific competition
We hypothesise that TH is dominated by a subset of the population that shows above-average dominance and aggression
Less dominant sharks are likely chased away or avoid close interactions with larger and dominant sharks
juvenile females generally avoided the feeding area when adult females were present
these behaviours are variable and require further investigation
our analyses show the residency to the dive site and not to the waters surrounding Fuvahmulah
It is likely that a large proportion of the population was present in the area but avoided the dive site and thus
we recommend incorporating all stakeholder’s interests into local management plans that support sustainable ecotourism in one of the world’s largest shark sanctuaries
Additional footage of tiger sharks was retrieved during exploratory usage of a remote underwater video station deployed by the Manta Trust and FDS (n = 9 encounters)
we calculated the mean to designate sharks in this study accordingly
female sharks that were estimated > 3 m were considered as adults and females estimated ≤ 3 m were considered juveniles
All tiger sharks were assigned their life stage after size estimates from 2022
If the individuals were not present in 2022
this approach has not been applied to sharks yet
Since our data set included photographs of the same sharks throughout their presumed gestation period and subsequent return after parturition
we used this method to quantify the observed morphological changes over time
To develop a dimensionless and scale-invariant metric for comparisons between photographs
we standardised the body widths by a length measure
we used the shark’s width from the posterior end of the first dorsal fin vertically down (90° to swimming direction) and the length from the anterior base of the pectoral fin to the anterior base of the pelvic fin to minimise error due to the sharks’ propulsive tail flexing (Supplementary Fig
where the picture quality allowed for the quantification of shark widths by having at least ten sufficient pictures spread over at least six months across presumed pregnancies (supplementary Fig
We calculated the ratios starting one year prior to a shark leaving for apparent parturition and took three measurements after its subsequent return
Sharks that did not fall into the category ‘pregnant’ or ‘not pregnant’
To assess if there was a consistent trend in abdominal distension throughout presumed gestation
we applied a linear regression model to the width data until the sharks left
The study was conducted following the guidelines and under the research permits issued by the Environmental Protection Agency (annually renewable permit: EPA/2021/PA-F01 and EPA/2022/PA-F02) and the Ministry of Fisheries
Maldives (annually renewable permit: 30-D/PRIV/2021/190)
The methods employed were non-invasive in nature
ensuring no harm was caused to the animals involved
All methods used were in accordance with ARRIVE guidelines
All data is available upon reasonable request to the corresponding author
Extinction risk and conservation of the world’s sharks and rays
Status and ecological effects of the world’s largest carnivores
Half a century of global decline in oceanic sharks and rays
Overfishing drives over one-third of all sharks and rays toward a global extinction crisis
Predicting ecological consequences of marine top predator declines
Cascading top-down effects of changing oceanic predator abundances
Patterns and ecosystem consequences of shark declines in the ocean
Global economic value of shark ecotourism: implications for conservation
Evidence of increased economic benefits from shark-diving tourism in the Maldives
State of the global market for shark products
Management of shark fisheries in the Maldives
Government of the Maldives. The President’s Office. Government to impose ban on trade and export of sharks and shark products. The President’s Office Republic of Maldives (2010). https://presidency.gov.mv/Press/Article/998
A global overview of shark sanctuary regulations and their impact on shark fisheries
The impact of injury on apparent survival of whale sharks (Rhincodon typus) in South Ari Atoll Marine Protected Area
Environmental drivers of reef manta ray (Mobula alfredi) visitation patterns to key aggregation habitats in the Maldives
UNESCO & Fuvahmulah Biosphere Reserve. UNESCO (2020). https://en.unesco.org/biosphere/aspac/fuvahmulah
The IUCN Red List of Threatened Species 2019: e.T39378A2913541 (2019)
Sharks caught in the KwaZulu-Natal bather protection programme
sex and reproductive biology of seven pelagic sharks in the Eastern Arabian Sea
New insights into the reproductive biology of the tiger shark Galeocerdo cuvier and no detection of polyandry in Reunion Island
Declining trends in annual catch rates of the tiger shark (Galeocerdo cuvier) in Queensland
Observations on the reproductive cycles of some viviparous North American sharks
A perspective on future tiger shark research
Genetic population structure and demography of an apex predator
Diagnosing the dangerous demography of manta rays using life history theory
Algumas considerações sobre a reprodução do cação jaguara
Reproductive biology of the tiger shark (Galeocerdo cuvier) in Hawaii
Global population genetic dynamics of a highly migratory
Population structure and connectivity of tiger sharks (Galeocerdo cuvier) across the Indo-Pacific Ocean basin
Reef-fidelity and migration of tiger sharks
Retrospective genomics highlights changes in genetic composition of tiger sharks (Galeocerdo cuvier) and potential loss of a South-Eastern Australia population
Exploring intraspecific life history patterns in sharks
Habitat geography around Hawaii’s oceanic islands influences tiger shark (Galeocerdo cuvier) spatial behaviour and shark bite risk at ocean recreation sites
Telemetry and random-walk models reveal complex patterns of partial migration in a large marine predator
Tiger shark (Galeocerdo cuvier) movement patterns and habitat use determined by satellite tagging in Eastern Australian waters
Long-Term movements of tiger sharks satellite-tagged in Shark Bay
Refuges and risks: evaluating the benefits of an expanded MPA network for mobile apex predators
Persistent transboundary movements of threatened sharks highlight the importance of cooperative management for effective conservation
Residency and movement patterns of an apex predatory shark (Galeocerdo cuvier) at the Galapagos Marine Reserve
A matter of taste: spatial and ontogenetic variations on the trophic ecology of the tiger shark at the Galapagos Marine Reserve
Seasonal variability of bull and tiger shark presence on the West coast of Reunion Island
long-distance migrations by a philopatric predator targeting highly contrasting ecosystems
Seasonal and life-stage variation in the reproductive ecology of a marine apex predator
Heeding the History of Kahu Mano: Developing and Validating a Pono Photo-Identification Methodology for Tiger Sharks (Galeocerdo cuvier) in Hawai’i (University of Hawai’i at Hilo
In shark research: emerging technologies and applications for the field and laboratory (ed
Annual re-sightings of photographically identified white sharks (Carcharodon carcharias) at an Eastern Pacific aggregation site (Guadalupe Island
Population density of striped hyenas in relation to habitat in a semi-arid landscape
residency patterns and movements of whale sharks in Southern Leyte
Philippines: results from dedicated photo-ID and citizen science: Whale sharks of Southern Leyte
behavior and demographics of reef manta rays in Komodo National Park
Analysis of animal movement using opportunistic individual identifications: application to sperm whales
In-water methods reveal population dynamics of a green turtle Chelonia mydas foraging aggregation in the Philippines
Using citizen science in the photo-identification of adult individuals of an amphibian based on two facial skin features
Pupping areas and mortality rates of young tiger sharks Galeocerdo cuvier in the Western North Atlantic Ocean
and regional movement of tiger sharks (Galeocerdo cuvier) at a pupping location in the Bahamas
Acoustic telemetry validates a citizen science approach for monitoring sharks on coral reefs
Photo-ID and telemetry highlight a global whale shark hotspot in Palawan
Satellite tagging and photographic identification reveal connectivity between two UNESCO World Heritage Areas for Reef Manta Rays
Behavioural patterns of a tiger Shark (Galeocerdo cuvier) feeding aggregation at a blue whale carcass in Prony Bay
persistence and impacts of residual fishing hooks on tiger sharks
Review of Shark Diving Practices at Fuvahmulah
Maldives72 (Prepared for the Ocean Country Partnership Programme
Growth and maximum size of tiger sharks (Galeocerdo cuvier) in Hawaii
Emmons, S. M., D’Alberto, B. M., Smart, J. J. & Simpfendorfer, C. A. Age and growth of tiger shark (Galeocerdo cuvier) from Western Australia. Mar. Freshw. Res.https://doi.org/10.1071/MF20291 (2021)
Don’t bite the hand that feeds: assessing ecological impacts of provisioning ecotourism on an apex marine predator: ecological impacts of shark ecotourism
& Species catalogue sharks of the world
An annotated and illustrated catalogue of shark species known to date
Fuvahmulah Atoll ISRA Factsheet (IUCN SSC Shark Specialist Group
Ocean acidification and global warming impair shark hunting behaviour and growth
Hand feeding can periodically fuel a major portion of bull shark energy requirements at a provisioning site in Fiji
Effects of food provisioning on the daily ration and dive site use of great hammerhead sharks
Shark depredation in commercial and recreational fisheries
localised movements and behavioural segregation of pregnant Carcharias taurus at Wolf Rock
An elasmobranch maternity ward: female round stingrays Urobatis halleri use warm
restored estuarine habitat during gestation
Aggregation behavior and seasonal philopatry in male and female leopard sharks Triakis semifasciata along the open coast of southern California
regional trends and inshore environmental conditions influence coral bleaching in Hawaii: CORAL BLEACHING IN HAWAII
Concurrent scavenging off a whale carcass by great white sharks
Life after death: behaviour of multiple shark species scavenging a whale carcass
Social preferences of juvenile lemon sharks
Social network analysis reveals potential fission-fusion behavior in a shark
Companions and casual acquaintances: the Nature of associations among bull sharks at a shark feeding site in Fiji
Social network analysis reveals the subtle impacts of tourist provisioning on the social behavior of a generalist marine apex predator
Aquatic animal telemetry: a panoramic window into the underwater world
A novel intrauterine satellite transmitter to identify parturition in large sharks
and research priorities of shark diving tourism
The shark reef marine reserve: a marine tourism project in Fiji involving local communities
Grey Nurse Shark (Carcharias taurus) diving tourism: Tourist compliance and shark behaviour at Fish Rock
Multi-year effects of wildlife tourism on shark residency and implications for management
Measuring whale sharks Rhincodon typus with laser photogrammetry
Long-term photo-identification reveals the population dynamics and strong site fidelity of adult whale sharks to the coastal waters of Donsol
Eyes on the size: accuracy of visual length estimates of white sharks
Paired-laser photogrammetry as a simple and accurate system for measuring the body size of free-ranging manta rays Manta alfredi
Risks and advantages of using surface laser photogrammetry on free-ranging marine organisms: a case study on white sharks Carcharodon carcharias
First record of a potential neonate tiger shark (Galeocerdo cuvier) at a remote oceanic island in the Eastern Tropical Pacific
Underwater ultrasonography and blood sampling provide the first observations of reproductive biology in free-swimming whale sharks
Spy in the sky: a method to identify pregnant small cetaceans
Assessment of a non-invasive approach to pregnancy diagnosis in gray whales through drone-based photogrammetry and faecal hormone analysis
SOCPROG programs: analysing animal social structures
Population structure and residency of whale sharks Rhincodon typus at Utila
Improving sightings-derived residency estimation for whale shark aggregations: a novel metric applied to a global data set
Selection of models of lagged identification rates and lagged association rates using AIC and QAIC
R Core Team. R: A language and environment for statistical computing. R Foundation for Statistical Computing, Vienna, Austria, URL (2022). https://www.R-project.org/
Download references
This work was possible through the support of Fuvahmulah Dive School and dedicated staff members (including A
Rasheedh) collecting video footage of tiger sharks across the seasons
Data collection was also supported by a wide range of citizen scientists and K
Zerr (Manta Trust) sharing their footage with us
Müller for additional help with study design
Open Access funding enabled and organized by Projekt DEAL
Leibniz Centre for Tropical Marine Research
Marine Research and Conservation Foundation
Department of Biological and Environmental Sciences
TD and TI designed and conceived the project
LV prepared the manuscript and data analysis
JC and LV collected data during the period of highest sampling effort from 2021 until 2023
MC designed the original photo identification catalogue and collected data in 2019 and 2020
IS and TI collected data throughout the entire study period
AN filmed the predation event on a hawksbill turtle
All authors contributed to manuscript revision
The authors declare no competing interests
Springer Nature remains neutral with regard to jurisdictional claims in published maps and institutional affiliations
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but a population in the island paradise could hold clues to where they breed and give birth
with their characteristic stripes and sharp
appear from the depths of the Indian Ocean
They follow the scent of fish blood and oil coming from tuna heads a research team has hidden under a pile of rocks in a shallow sandy area
perhaps eight or nine and up to four metres long
two with bite marks on their flanks and fins
encounters with more than 20 sharks are not uncommon
Tiger sharks circle at the Tiger Zoo dive site
Earlier this year, scientists, two local dive schools and the Maldivian shark organisation Miyaru teamed up for a two-week field study to answer the question: why is it that so many tiger sharks gather near the island
The findings could help locate what is considered to be the holy grail of shark research: the place where the sharks go to breed and life begins
we need to know the habitats that are critical for their reproduction,” says Lennart Vossgaetter
a German shark research and expedition company
these places are mostly unknown for migratory species such as tiger sharks
which roam the oceans over thousands of kilometres
he continued what a local dive school had started: documenting which sharks showed up on the dives using photo-IDs
A dive guide hides tuna heads under a pile of rocks at the site to attract the sharks
we need to know the habitats that are critical for their reproduction,’ he says
With the help of three other biologists carrying out identifications
“making it the largest known population of tiger sharks in the ocean”
He noticed that many got fatter and fatter over time – only to disappear for months and return with flat bellies: did the tiger sharks spend their pregnancy in these waters
This would make it a key region for the species’ survival in the Indian Ocean
the research team enlisted the help of James Sulikowski
He co-leads the Big Fish Lab at Oregon State University and studies shark migration and reproduction
It is Sulikowski and his team who planted the tuna heads as bait to enable them to carry out an important but delicate procedure
Sulikowski then reaches out with a telescopic pole and presses an ultrasound probe against the shark’s belly
James Sulikowski and the team examine sharks for pregnancy with the underwater ultrasound device
we need to protect their critical habitats and migration routesLennart VossgaetterBack at the dive club
Sulikowksi displays an ultrasound scan of a shark named Aaya: she is indeed pregnant
their gills and pectorals are clearly visible
“Aaya carries about 40 of them.” When she gives birth after about 16 months – nobody knows for sure how long the gestation is – they will have reached 75cm
the researchers scan 35 female tiger sharks
Why the females visit this particular region during their pregnancy is unclear
“Perhaps because they are safe here from aggressive males,” Sulikowski says
“They could also seek out this warm region to accelerate the growth of the embryos
as their metabolism depends on the water temperature.”
Whatever the reason, it is good news for tiger sharks that one of their reproduction sites in the Indian Ocean, a hotspot of global shark mortality, falls within the protected area of the Maldives
Clockwise: a boat at the entrance to the harbour dive site; a beach popular with locals and tourists at the northern end of the island; and fishermen selling tuna
In 2017, Tatiana Ivanova and her husband opened the first dive centre on the island, Fuvahmulah Dive School, and sparked a boom in the trade. Now there are about a dozen such centres. The reef-fringed island offers high chances to see big fish such as manta rays, thresher sharks and hammerheads. But its main attraction is the tiger sharks – at Tiger Zoo, sightings are guaranteed. “Ninety-five per cent of our guests come for them,” says Ivanova.
Tatiana Ivanova and Ibrahim Siyan, owners of Fuvahmulah Dive School
Read moreWhere they mate and give birth the team can still only speculate. “Our guess is that pregnant females visit another atoll in the Maldives,” says Vossgaetter. But they could also swim to the Chagos archipelago, a marine reserve where sharks are caught illegally, or to Sri Lanka, where they are not protected at all.
In autumn, the researchers plan to return to Fuvahmulah to follow the progress of the tiger sharks’ pregnancies. In the next phase, they would like to tag females using a newly developed “birth alert tag”, a chicken egg-sized transmitter that is inserted into the uterus. At birth, the tag is released together with the baby sharks and transmits its position via satellite. If successful, it would help better understand the life of tiger sharks and protect this apex predator.
Volume 7 - 2020 | https://doi.org/10.3389/fmars.2020.538675
Assessing the resilience of islands toward altered ocean climate pressures and providing robust adaptation measures requires an understanding of the interaction between morphological processes and the underlying hydrodynamic drivers
this study presents changing sediment volumes on various temporal scales for the fringing reef island Fuvahmulah
aerial imagery provides information on marine aggregates of the island's beaches
high resolution climate reanalysis data serves as input into an empirical and a numerical approach
both approaches describe the driving processes behind volumetric seasonal and interannual changes: On the one hand
the empirical method quantifies sediment transport rates for calcareous sediments over the whole time span of the data set by considering wind and swell waves from multiple directions
the numerical method gives insights into the complexity of currents induced by dominant wave components
Combining these methods facilitates hindcasting and predicting morphological changes under varying wave climate
assessing sediment pathways over the whole reef
and describing the seasonal and interannual evolution of the sand spit Thoondu
this study reveals sediment distribution on different spatio-temporal scales and elucidates their significance in the design of conventional and alternative low-regret coastal adaptation
This shows that sediment erosion and accretion is highly site specific and is sensitive to multiple drivers
the evolution of reef islands and their natural response to changing impacts mainly depend on the reef's individual sediment production rate and the sediment distribution under the given hydrodynamic circumstances:
monsoon-driven seasonal as well as the annual variations of the local hydro- and morphodynamic regime
numerical modeling efforts help to further analyze coral sediment entrainment
transport processes and distribution on the reef and around the island
dependent on wave directionality and forcing
These approaches mimic the natural seasonal and annual variations of the typical local hydro- and morphodynamic regime
Considering multi-directional wave input facilitates evaluating characteristic regional and local wave climate as well as following the probable sediment pathway around the island
Compared to studies on annual or seasonal scale
the hourly temporal resolution of hydrodynamic data gives new and more detailed insights into drivers of morphodynamics on reef islands and thus improves the understanding of sediment distribution
scrutinizes the influence of a varying wave climate on reef sediment pathways
and thereby showcases reef islands' capability to naturally cope with changing ocean and climate pressures on reef islands
this potential must be considered in coastal management and improves conventional and alternative coastal protection measures for sustainable low-regret coastal adaptation
Cartopy also delivered the coastlines in (B,C)
Figure 2. Wave climate at the ERA5 data node (Figure 1) (A) Shows monthly box plots of hourly significant wave height Hs for a time series from 1979 to 2019
blue boxes are wet season months and yellow boxes are dry season months
(B) Shows the discrete marginal distribution between the peak wave direction θp and the significant wave height Hs of the dry season
while (C) contains this information for the wet season
while (D,E) contain the same information for wind waves
the colorbar shows the occurrence probability of each 0.5° wave direction and 0.05 m wave height cell
The gray areas visualize the significant peak direction range θp
containing the directions which combined have the highest 33 % occurrence probability
Table 1. Wave parameters for the time series at the ERA5 data node (Figure 1)
Three field campaigns provided data to evaluate the seasonal and annual sedimentation processes around the island: The first campaign started in March 2017 at the end of the dry season
The second field campaign took place at the end of the wet season in September 2017
revealing seasonal variations in combination with the former measurements in March
The last field measures were taken in March and April 2019
disclosing seasonal and annual changes of sediment volumes when compared to the data recorded in 2017
The first field campaign also accommodated a bathymetric survey
recording depth profiles with a dual-frequency echo-sounder (Dr
the field campaigns yielded the following data sets:
Quantum GIS (version 3.4.11.) uses the rastered topography provided by Photoscan to calculate the seasonal or annual changes of the coastal topography with
resulting in a difference of topographic height Δzi, j for each pixel of the rastered DEM. All further post-processing (Figure 3) was performed with the Python module Rasterio (version 1.1.1): The total volume change in each of the ΔDEM is calculated with
ΔVmc equals to ΔVDEM but only within the boundaries of the morphological cell (index mc)
Summarizing the areal sediment volumes of each morphological cell for the entire island with the absolute areal sediment difference quantifies the morphodynamic activity and gives an estimate on the average shoreline change (index sc):
Average areal sediment budgets from DEMs with (A) seasonal sediment budget from dry to wet season
based on the data of March 2017 and September 2017 and (B) annual sediment budget between dry season 2017 and 2019 (both recorded in March)
sediment accumulation for the observation period is blue
The volumetric change of sediment [m3] is averaged over dry beach area [m2] (see Equation 4)
dashed) come from measurements in the first field campaign (March 2017)
Location of sediment volumes and profiles (orange lines) for the dry season 2017 (red colorbar) and wet season 2017 (blue colorbar) in the north of Fuvahmulah
In the dry season sand depots form at the adjacent beaches Geiymiskih Fannu in the west (profile 1) and Thoondu in the east (profile 3) of the headland
The wet season allocates sand on the north tip of the island
The dry season sand volume of eastern Thoondu beach (profile 3) was integrated over differences of six accuracy-improved profiles
The profile elevation N is given as distance to the WGS84 ellipsoid
The background satellite image contains Copernicus Sentinel data (true color
large areas of the reef were covered with sand in the dry season
making vGCP assignment on the sea-side of the beach impossible
so that DEMs contain areas with either poor or without referencing
six beach profiles support the estimation of sand volumes on the dry season Thoondu beach
The profiles have been manually referenced to the plain beach in wet season
provided by ©OpenStreetMap contributors
Combining the annual areal carbonate production rate with the reef platform gives the total annual estimate of sediment volume produced on the reef as
with a measured sediment density of ρs= 2 g cm−3
The CERC-formulation or Shore Protection Manual (SPM)-method for longshore sediment transport (CERC, 1984) uses the longshore component of wave power as longshore wave energy flux
Converting Equation (7) to deep water parameters with n0 = ½ yields:
here with salt water density ρw of 1.03 g cm−3
sediment density ρs of 2 g cm−3
The empirical coefficient K is the transmission between wave-induced energy flux and sediment parameters. It accounts for the heterogeneity of beach sediments and the local coastal environments. The coefficient translates the wave energy impact on site into the site-specific sediment motion by calibration with measured data (Schoonees and Theron, 1995)
K facilitates matching the measured with the calculated total sediment volume of the field campaigns in 2019
the total sediment volume transported on a coast for all wave partitions is
with wpww for wind waves and wpSi for swell waves with i for each of the three swell partitions of the ERA5 data set
With the sediment volumes measured in the 2019 dry season field campaign for the northwest (NW: 14.689 m3) and east coast (E: 1.171 m3)
the empirical coefficient K becomes KNW = 55.89 and KE = 107.26
When illustrating results from the CERC method based on the hourly ERA5 data between January 2016 and November 2019
this study smooths the data with a combined windowed forward and backward average filter based on
with a window of n = 24 h·7 d. Filtering was applied on Hs (depicted in Equation 12), θp, qy, and Qy (analogous to Equation 12). Smoothing accounts for a delayed response of the morphology to wave energy forcing (Rutten et al., 2017)
Fuvahmulah has two seasons: the wet or rainy season from May to October and the dry season from November to April. Between the wet and the dry season, waves differ in height as shown by the ERA5 data set (C3S, 2017) of the ECMWF (Figure 2). While the dry season brings northeastern winds, Fuvahmulah is subject to mainly south to southwestern Monsoons in the wet season (MEE, 2014)
The Monsoons are accompanied by high waves with the highest median significant wave height in July (H~s,max.=1.72m)
while the highest total significant wave height was registered on November
While this study defines November already as dry season
the wave data discloses a transition phase between wet and dry season
The lowest annual median value H~s,min.=1.12m appears in March (dry season)
These parameters are used to describe the dominant features of wave heights
Around Fuvahmulah, total swell has a higher seasonal variation of Hs but less spreading or directional variety of associated peak wave directions θp than wind waves (Figure 2 and Table 1) Swell waves are rarely below 1 m in the wet season (Figure 2
ts are coming from south to southwest in both seasons
however the dry season contains additional directional components between ~45 and 135°
Wind waves have a mean significant height of H¯s,ww=0.26±0.28m (in both seasons, Figure 2 and Table 1)
The highest wind waves approach the island from west
The two significant differences among seasonal wind waves are (a) in the dry season waves reach Fuvahmulah dominantly between 238 and 75° while they approach the island with 122–287° in the wet season and (b) an additional north to west (0–90°) component in the dry season being absent in the wet season
Both differences have a considerable influence on the sediment transport in the northern part of the island and the sand spit Thoondu
The Gnaviyani Atoll reef is mostly sub-tidal with an inter-tidal reef-flat around Fuvahmulah
The sub-tidal reef fringes the inter-tidal plateau before it expands to an about ~2.2 km long and 1.6–0.5 km wide fore reef in the south east
This southern fore reef makes about 58.8 % of the total reef area
the depth is ~ 25 m and slowly decreases to around 8–15 m until reaching the inter-tidal reef
The reef form is elliptical with the long axis approximate to the dominant swell wave direction θp ~ 157.5°
The transition from sub-tidal fore reef to intertidal reef on the south and west side is sudden
while it is more gradual on the north and east side of Fuvahmulah
The inter-tidal reef is wider on the south and west side (around 85–145 m) than on the north and east side (around 15–25 m)
The island oscillation index Io describes the morphodynamic activity of an entire island by comparing the island's perimeter with the coastal area encompassing more than half of the maximum (seasonal) coastline change
The coastline change is linked to the beaches' volumetric changes and—in this study—estimated by integrating seasonal and interannual sediment differences over 12.5 m wide areal increments of Fuvahmulah's coast
The underlying sediment differences required for this procedure originate from the DEMs of the UAV surveys
Using this information for the island oscillation index leads to a low Io of Io = 350° − 315°/360° = 9.7 %. Such low oscillation indices on elliptical islands indicate that the elongated, lateral shorelines are meant to be morphologically stable (Kench and Brander, 2006) if not obstructed by artificial interference (Kench, 2012)
Focusing on the prominent morphodynamic area in the north of Fuvahmulah discloses the seasonal formation of the Thoondu spit:
The UAV-borne data of the two field campaigns in 2017 discloses a positive seasonal sediment balance
The instantaneous records do not contain the maximum depot sizes
as the seasonal difference in sediment volume is +12.307 m3
In the field campaign during the dry season of 2019
sediment depots formed at the same locations as in the dry season 2017
the depot in the west (Geiymiskih Fannu) was considerably larger when compared to 2017 (14.689 m3)
while the eastern depot was smaller (1.171 m3
the sediment volume recorded in the dry season of 2019 is 15.860 m3
This is an additional sediment volume of +6.361 m3 over the dry season of 2017
while it is -5.946 m3 less than in the wet season of 2017
the empirical and numerical methods can capture the spatio-temporal sediment transport over the reef
(B) Current velocities from the depth-averaged
phase resolving numerical wave model (BOSZ) for waves approaching the island from θp = 135°
(C) θp = 157.5° and (D) θp = 202°
The background color shows the magnitude of wave induced current velocity on the reef while the arrows depict the local current direction
Dashed red contour lines are the reef depths −5 and -15 m recorded in the first field campaign (March 2017)
Table 2. Beach orientation as well as longshore sediment transport directions and energy flux rates Py, i from the CERC-formulation (Figures 5A, 7, 8)
max and the contribution of each wave partition to Qy)
The time series shows values from January 2016 to November 2019
while negative values lead to sediment transport toward Thoondu beach in the northeast
The time series shows the dry season with orange and the wet season with blue background
Field campaign 1 and 3 (orange hatched area) took place in March 2017 and 2019
Field campaign 2 (blue hatched area) was conducted in September 2017
Table 3. Peak transport rates qy, max and the contribution of each wave partition to the total transported sediment volume Qy in the CERC-formulation between 2016 and 2019 on the northern beaches Geiymiskih and Thoondu (NW: Figure 7 and E: Figure 8)
The resulting dominant longshore sediment transport direction is toward Thoondu beach (67.9 %)
The dominant sediment transport toward Thoondu is commonly found in the course of the wet season
usually peaking between July and early September
Southward components only appear in dry season
wp indicate a transport toward the seaport in the southeast
while positive values lead to sediment transport toward Thoondu beach in the northwest
After the Thoondu spit formed in the wet season
the dry season wave climate leads to sediment depletion of the Thoondu spit
while new sediment accumulations form on Geiymiskih and Thoondu beach
Figure 9. Sentinel satellite images of northern Fuvahmulah on October 9, 2016 (A) and November 28, 2016 (B) and longshore sediment transport on Thoondu (C) and Geiymiskih Fannu beach (D) at the associated time of the satellite images (green line). (C,D) Show zoomed data from the longshore sediment transport rates qy, wp (Figures 7, 8)
The orange hatched area shows the time of the first field campaign for this study
The satellite images contain Copernicus Sentinel data
Sediment distribution on a reef is subject to (a) hydrodynamic forcing
defining wave propagation and thus current direction on the reef platform
as well as (c) sediment supply of the reef
This study used wave data from the climate reanalysis model ERA5 in the process-based
empirical CERC method and the phase-resolving
depth-integrated numerical model BOSZ to study the associated morphodynamic response on the Maldivian reef island Fuvahmulah
the study linked these insights to comprehend measured shoreline changes on the island's beaches and the seasonal formation of the sand spit Thoondu in the north of the island
These factors—the good spatio-temporal data availability and trustworthy reconstruction of the wave climate—facilitate event based hind- and forecasting in the CERC formulation
the sheer amount of wave data results in a challenging number of feasible and reasonable numerical simulations
this study combines the empirical CERC-formulation with information of a numerical model
calculating wave-induced currents approaching Fuvahmulah from the dominant wave directions:
The sediment transport rate qy itself focuses on longshore sediment distribution but gives no information on cross-shore beach response (Atkinson et al., 2018) or cross-shore transport components
the CERC-approach cannot estimate sediment transport toward the island and thus
the role of the southern Fuvahmulah reef as sediment source
The numerical models also disclose an influence of south to southeastern swells on the lee-side of the island
but these are outside the northern beaches' incident wave direction range θ⊥ of −90–90° and thus remain unconsidered in the CERC-equation
The insights from the numerical model complement the findings of the CERC-method and put the measured morphological changes into an hydrodynamic context
erosion must be studied on a different scale
Ocean climate pressures influence sediment production and distribution pathways on the reef
The dynamic reef island morphology is a natural response to changing impacts
low-regret coastal development requires an understanding of the underlying processes of island morphology
Morphological changes on the reef are triggered by hydrodynamic forcing. Therefore, this study exploits hourly climate reanalysis model wave data to quantify longshore sediment distribution around the island with the CERC method. Dominant south to southeasterly swells are characteristic for the regional Maldivian hydrodynamic regime (Kench and Brander, 2006; Kench and Mann, 2017)
These swells induce a constant sediment supply from the southeastern reef plateau
The sediment stream goes around the lateral sides of Fuvahmulah to the northern headland
where the Thoondu spit forms in the wet season
Local wind-wave and subordinate swell events shape the northern beaches within the seasons
By adapting and validating the empirical coefficient K with the measurements from the field campaigns
the CERC formulation is capable of capturing morphodynamic changes after hydrodynamic events
This method facilitates processing the high resolution wave data from the climate data models for quantitative hind- and forecast of longshore sediment transport
Numerical methods give supplementary insights beyond the empirical CERC approach and hence a more holistic view on the complex hydrodynamic background of the reef for certain combinations of dominant wave parameters or single events
further advances in numerical models are required to directly couple hydrodynamic drivers and morphodynamic response on reef islands and calcareous sediment
sediment trajectories reveal locations with increased erosion potential on the lee-side of disturbances in the natural sediment stream
Because anthropogenic interventions into the natural dynamics disturb the natural ability to respond to ocean climate pressures the design of coastal infrastructure on low-lying reef islands must include an assessment of reef islands' natural beach nourishment potential to avoid interference before taking action
reef island communities can reduce shoreline erosion
improve their resilience and decrease their risk to suffer from sea level rise and associated impacts
The field data supporting the findings of this study are available from the corresponding author upon reasonable request. Wave climate data is available from the Copernicus Climate Change Service (C3S, 2017)
Satellite images from Copernicus Sentinel data were accessed through the EO Browser
and wrote the manuscript with input from TS
All authors contributed to the article and approved the submitted version
This study took place in the project Dealing with change in SIDS: societal action and political reaction in sea level change adaptation in Small Island Developing States (DICES), grant no. SCHL 503/17-1. The project was framed within the priority program (SPP 1889) ‘Regional Sea Level Change and Society’ (SeaLevel, see www.spp-sealevel.de) of the German Research Foundation (Deutsche Forschungsgemeinschaft
The publication of this article was funded by the Open Access fund of Leibniz Universität Hannover
The authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest
and Uwe Zander for their help in the field campaigns
the authors were supported by the Maldives Meteorological Service (MMS)
The authors further appreciate the involvement of the working group of Maike Paul for reviewing the manuscript
We further acknowledge the working group of Jan Visscher
as well as our project members in DICES Susann Adloff
and Katrin Rehdanz for their professional input
Additionally the authors thank Volker Roeber and Jannek Gundlach for their feedback concerning numerical models as well as Jean Bidlot
and Todd Spindler for their help in accessing climate reanalysis data
Elisa Casella and Alessio Rovere gave valuable insights and feedback to UAV-based photogrammetry
while Tobias Kersten helped with geodetic questions before the third field campaign and while post-processing
The Supplementary Material for this article can be found online at: https://www.frontiersin.org/articles/10.3389/fmars.2020.538675/full#supplementary-material
A fundamental paradigm for coral reef carbonate sediment dissolution
CrossRef Full Text | Google Scholar
Laboratory investigation of the bruun rule and beach response to sea level rise
Swash–groundwater interaction on a steep gravel beach
CrossRef Full Text | Google Scholar
Reading: European Centre for Medium-RangeWeather Forecasts (ECMWF)
Google Scholar
Google Scholar
“Design tools related to engineering,” in Environmental Design Guidelines for Low Crested Coastal Structures
C3S (2017). ERA5, Fifth Generation of ECMWF Atmospheric Reanalyses of the Global Climate by the Copernicus Climate Change Service (C3S). Copernicus Climate Change Service Climate Data Store (CDS). Available online at: https://cds.climate.copernicus.eu/cdsapp#!/home (accessed December 6
PubMed Abstract | Google Scholar
Increased frequency of extreme Indian Ocean Dipole events due to greenhouse warming
Sea level Measured by Tide Gauges From Global Oceans-The Joint Archive for Sea Level Holdings (NCEI Accession 0019568)
NOAA National Centers for Environmental Information
Google Scholar
CrossRef Full Text | Google Scholar
Mapping coral reefs using consumer-grade drones and structure from motion photogrammetry techniques
Accuracy of sand beach topography surveying by drones and photogrammetry
CrossRef Full Text | Google Scholar
MS: Coastal Engineering Research Center (CERC)
Google Scholar
CrossRef Full Text | Google Scholar
Observations of wave transformation over a fringing coral reef and the importance of low-frequency waves and offshore water levels to runup
abrupt changes managing risk,” in IPCC Special Report on the Ocean Cryosphere in a Changing Climate
Google Scholar
Persistent acceleration in global sea-level rise since the 1960s
Generation and propagation of ship-borne waves–solutions from a Boussinesq-type model
CrossRef Full Text | Google Scholar
“Coastal Infrastructure on Reef Islands–the Port of Fuvahmulah
the Maldives as example of maladaptation to sea-level rise?” in Coastal Structures 2019
Schlurmann (Hannover: Bundesanstalt fr Wasserbau)
Google Scholar
“Assessing the application potential of selected ecosystem-based
low-regret coastal protection measures,” in Ecosystem-Based Disaster Risk Reduction and Adaptation in Practice
Nehren (Cham: Springer International Publishing)
Rapid human-driven undermining of atoll island capacity to adjust to ocean climate-related pressures
CrossRef Full Text | Google Scholar
Coral reef island initiation and development under higher than present sea levels
doi: 10.1175/1520-0469(1988)045<1143:TMIOH>2.0.CO;2
CrossRef Full Text | Google Scholar
The effectiveness of coral reefs for coastal hazard risk reduction and adaptation
Identification and classification of very low frequency waves on a coral reef flat
Wave-generated flow on coral reefs—an analysis for two-dimensional horizontal reef-tops with steep faces
CrossRef Full Text | Google Scholar
Estimating regional coral reef calcium carbonate production from remotely sensed seafloor maps
Automated analysis of ocean surface directional wave spectra
doi: 10.1175/1520-0426(2001)018<0277:AAOOSD>2.0.CO;2
CrossRef Full Text | Google Scholar
Coral reef structural complexity provides important coastal protection from waves under rising sea levels
A sustained ocean observing system in the Indian Ocean for climate related scientific knowledge and societal needs
Wind-wave misalignment and a combination method for direction-dependent extreme incidents
CrossRef Full Text | Google Scholar
Performance benchmarking tsunami models for NTHMP's inundation mapping activities
“Managing the risks of extreme events and disasters to advance climate change adaptation,” in A Special Report of Working Groups I and II of the Intergovernmental Panel on Climate Change (IPCC)
Google Scholar
“Compromising Reef Island shoreline dynamics: legacies of the engineering paradigm in the Maldives,” in Pitfalls of Shoreline Stabilization: Selected Case Studies
CrossRef Full Text | Google Scholar
Response of reef island shorelines to seasonal climate oscillations: South Maalhosmadulu atoll
CrossRef Full Text | Google Scholar
Reef Island evolution and dynamics: insights from the Indian and Pacific Oceans and perspectives for the Spermonde Archipelago
CrossRef Full Text | Google Scholar
Influence of natural climate variability on the extreme ocean surface wave heights over the Indian Ocean
CrossRef Full Text | Google Scholar
Longuet-Higgins
Longshore currents generated by obliquely incident sea waves: 2
CrossRef Full Text | Google Scholar
Inter-model analysis of tsunami-induced coastal currents
CrossRef Full Text | Google Scholar
“Integrative cross-chapter box on low-lying islands coasts,” in Special Report on Ocean Cryosphere in a Changing Climate
Google Scholar
Analytical modelling of wave refraction and convergence on coral reef platforms: implications for island formation and stability
CrossRef Full Text | Google Scholar
Physical and numerical modeling of infragravity wave generation and transformation on coral reef platforms
Modelling storm hydrodynamics on gravel beaches with XBeach-G
Destruction or persistence of coral atoll islands in the face of 20th and 21st century sea-level rise
CrossRef Full Text | Google Scholar
MEE (2014). Environment & Social Assessment & Management Framework–Climate Change Adaptation Project. Technical report, Ministry of Environment and Energy. Available online at: http://documents.worldbank.org/curated/en/939971468329099370/pdf/E47010SAR0EA0P1533010Box385392B00PUBLIC0.pdf (accessed December 27
Google Scholar
Möller
Wave attenuation over coastal salt marshes under storm surge conditions
Foahmulaku Beach Erosion Survey & Coastal Protection Report
Google Scholar
costs and coastal protection benefits of natural and nature-based defences
Infiltration effects on sediment mobility under waves
CrossRef Full Text | Google Scholar
“Small islands,” in Climate Change 2014
Contribution of Working Group II to the Fifth Assessment Report of the Intergovernmental Panel of Climate Change
“Sea Level rise implications for low lying islands
coasts communities,” in IPCC Special Report on the Ocean Cryosphere in a Changing Climate
Google Scholar
Let it flow: how does an underlying current affect wave propagation over a natural seagrass meadow
CrossRef Full Text | Google Scholar
Loss of coral reef growth capacity to track future increases in sea level
Implications of reef ecosystem change for the stability and maintenance of coral reef islands
Changes in the population and community structure of corals during recent disturbances (February 2016–October 2017) on Maldivian coral reefs
Spectral wave-driven sediment transport across a fringing reef
Sediment transport in the presence of large reef bottom roughness
Spatial variability of sediment transport processes over intratidal and subtidal timescales within a fringing coral reef system
Challenges for shared responsibility–political and social framing of coastal protection transformation in the Maldives
Destructive tsunami-like wave generated by surf beat over a coral reef during Typhoon Haiyan
Boussinesq-type model for energetic breaking waves in fringing reef environments
CrossRef Full Text | Google Scholar
Shock-capturing Boussinesq-type model for nearshore wave processes
CrossRef Full Text | Google Scholar
Observations on sandbar behaviour along a man-made curved coast
CrossRef Full Text | Google Scholar
Massive corals maintain a positive carbonate budget of a Maldivian upper reef platform despite major bleaching event
Possible impacts of Indian Ocean Dipole mode events on global climate
CrossRef Full Text | Google Scholar
A dipole mode in the tropical Indian Ocean
Accuracy and Applicability of the SPM Longshore Transport Formula
Google Scholar
Assessing morphologic controls on Atoll Island alongshore sediment transport gradients due to future sea-level rise
CrossRef Full Text | Google Scholar
The relentless march of mass coral bleaching: a global perspective of changing heat stress
Dependence of total longshore sediment transport rates on incident wave parameters and breaker type
CrossRef Full Text | Google Scholar
A new alternative to saving our beaches from sea-level rise: the sand engine
Many Atolls may be uninhabitable within decades due to climate change
Most atolls will be uninhabitable by the mid-21st century because of sea-level rise exacerbating wave-driven flooding
Ecosystem-based coastal defence in the face of global change
Profiling risk and sustainability in coastal deltas of the world
“Tropical cyclone formation,” in Global Perspectives on Tropical Cyclones (San Jose
CrossRef Full Text | Google Scholar
Swash infiltration-exfiltration and sediment transport
CrossRef Full Text | Google Scholar
Principles of Sedimentation and Erosion Engineering in Rivers
Google Scholar
CrossRef Full Text | Google Scholar
Synergy of wind wave model simulations and satellite observations during extreme events
Consumer-grade global positioning system (GPS) accuracy and reliability
CrossRef Full Text | Google Scholar
Atoll reef-island formation and response to sea-level change: West Island
CrossRef Full Text | Google Scholar
Citation: David CG and Schlurmann T (2020) Hydrodynamic Drivers and Morphological Responses on Small Coral Islands—The Thoondu Spit on Fuvahmulah
Received: 28 February 2020; Accepted: 29 September 2020; Published: 29 October 2020
Copyright © 2020 David and Schlurmann. This is an open-access article distributed under the terms of the Creative Commons Attribution License (CC BY)
distribution or reproduction in other forums is permitted
provided the original author(s) and the copyright owner(s) are credited and that the original publication in this journal is cited
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distribution or reproduction is permitted which does not comply with these terms
*Correspondence: C. Gabriel David, ZGF2aWRAbHVmaS51bmktaGFubm92ZXIuZGU=
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beyond the country’s legendary resorts
cruising between atolls on a luxury liveaboard
tasting regional food grown on local farms
observing goldsmiths crafting traditional necklaces
These may not be typical activities associated with the Maldives
a tropical archipelago best known for its high-end resorts
holiday experiences can find Maldivian guest houses or liveaboards that open doors to new perspectives and unlock access to new experiences
Perhaps one of the most thrilling of these is the bucket list-worthy experience of diving with huge aggregations of tiger sharks
“We have arguably one of the biggest tiger shark aggregations in the world,” says Fuvahmulah-based shark guide Hamna ‘Hamty’ Hussain
are always located on inhabited islands (unlike resorts
which are each on a private island of their own)
This offers you the unique opportunity to live alongside Maldivians and interact with the community
as well as support locally owned services such as dive centres
Liveaboard sailing adventures can also open opportunities to experience island life
chat with Maldivians and maybe even hear some mesmerising tales
There are shipwrecks all over the Maldives – one of the oldest dates back to the 1700s – and some of them
According to Maldivian oral history shared across generations
the Sultan of the Maldives’ goldsmith was banished from Malé to Rinbudhoo after being accused of stealing
Islanders say he was one of the best goldsmiths in the Maldives and passed down his skills to the people of Rinbudhoo
Islanders dove to retrieve gold from shipwrecks to use in their workshops and became the best goldsmiths in the Maldives
it was said that every house in Rinbudhoo had a goldsmith
you can still observe goldsmiths in action
As you wander around the streets of this peaceful island
stop by a tea shop to try some delicious hedika (Maldivian afternoon tea) with locals
made from Rinbudhoo’s unique curry blend of havaadhu spices
Due to the island’s tiny population (about 429)
the island has an atmosphere of tranquillity
with most of the inhabitants staying indoors under fans to avoid the heat of the day
only to emerge and socialise in the early morning and evenings
Running the length of the sleepy island are vibrant murals depicting Maldivian life
They are framed by an arch shaped like a golden fahtaru (gold necklace)
representing Rinbudhoo’s legendary goldsmiths
You can create your own island-hopping itinerary of local experiences in a cost-efficient way by utilising MTCC public ferry (dhoni) routes
Many guest houses or water-sports centres go beyond the typical offerings such as jet-skis and snorkelling
They bring you more uniquely Maldivian experiences such as sandbank picnics
traditional pole-and-line fishing trips at sunset and local cooking classes
save time by taking a domestic flight or consider a scheduled speedboat as an exhilarating alternative
Using high-speed scheduled routes such as the RTL ferry
a 500-year-old wooden palace in Utheemu island in the north of the country
Utheemu Ganduvaru was once the home of Sultan Mohamed Thakurufaanu who overthrew the Portuguese in 1573
It was a time when fishing villages and agricultural smallholdings were the norm
Just a couple of hops by local ferry from Rinbudhoo to South Ari Atoll is Dhigurah
a long shard of emerald green surrounded by azure waters
a guesthouse more akin to a stylish boutique hotel
join a liveaboard with a Central Atolls itinerary passing near (or docking at) Dhigurah
plus shared daily experiences such as diving and island visits
a sense of camaraderie typically develops fast among passengers
with the chance of making lifelong friendships
past the candy-coloured bungalows adorned with tropical flowers
Chat with a local relaxing on a joali (hammock chair) in the shade
watch boat-builders working on the beach and observe fishermen bringing their catches into the harbour
The fishing village is limited to just one end of this 3.2km-long
leaving more than 2.4km of untouched white sand
Another way to explore the Maldives is through its cuisine
Thoddoo in North Ari Atoll is a patchwork of green fields and orchards
located just 80 minutes by speedboat from Malé
and among fields bursting with juicy cucumbers and scotch bonnet chillies
Travellers can self-guide or book a farm tour through their guest house to learn about Thoddoo’s 112 farms
Maldivian chef Nasreena Ali is feted for promoting homegrown Maldivian cuisine
having shared her expertise at dining venues in Auckland and around the Maldives
She is currently a sous chef at Dhigali Maldives
“We use a lot of spices in traditional food
“The taste of Maldivian cuisine is different
Maldivian chillies and coconut milk.”
A visit to Thoddoo wouldn’t be complete without sampling its bounty at a farm-to-table eatery
Try the popular mas’huni and roshi (Maldivian breakfast of fresh tuna with onions
accompanied by a farm egg and hot sweet tea)
Or tuck into a lunch of grilled Maldivian octopus with a fresh salad of cucumber and peppers
the cuisine typically has strong ties to the ocean
but some islands like Thoddoo are very important islands for agriculture,” says Nasreena
“Thoddoo is particularly famous for its watermelons
or ‘Thoddoo karaa’ in Maldivian language.”
The jewel in the crown of the Maldives’ aquatic world is Baa Atoll Unesco World Biosphere Reserve
It’s home to globally significant biodiversity
the manta and whale shark “feeding frenzy” at Hanifaru Bay is the star attraction
This phenomenon sees extraordinary aggregations of manta rays attracted by zooplankton that get trapped in the horseshoe-shaped bay
Fayyaz Ibrahim comes from a family of divers
his father was the first Maldivian scuba diver and his uncle was the country’s first certified diving instructor
Fayyaz has worked on liveaboards and as an instructor for 42 years: “The mantas go into a feeding frenzy when they see the plankton
and sometimes on a good day you can see 200 there,” he says
only a licensed guide can take you for snorkelling
and there are special rules and rangers looking after the base
While Baa Atoll is an excellent location for snorkelling or learning to dive
the Maldives also offers thrilling experiences for more advanced divers
Fuvahmulah in the far south offers one of the Maldives’ most unforgettable experiences – diving with tiger sharks
Fuvahmulah’s only female shark guide Hamna Hussain
says: “Fuvahmulah is below the equator and it’s a one-island atoll
There are no other islands nearby and there’s no lagoon
It’s basically a pinnacle that just comes out of the deep water
we get all sorts of pelagic creatures like tiger sharks
we have some of the biggest aggregations of oceanic mantas
and one of the best coral reefs I’ve ever seen in the Maldives.” Tiger shark season runs from February to April in Fuvahmulah
These are just a handful of the diverse experiences to be discovered in the Maldives
liveaboards and islanders willing to share Maldivian culture with visitors
and year-round opportunities for adventure
people were coming during the European winter season,” says Fayyaz
“But now even in the European summer season people come
for the liveaboards and for my country.”
sand and sea of the sunny side of lifeBreathtaking views
Sitting just under the Equator at the southern extreme of the Maldives is the little-visited island of Fuvahmulah
It is very different from the hundred or so holiday resort islands dotted throughout the 800-kilometre chain of coral atolls stretching northwards in the Indian Ocean that make up the archipelago
This 4.5-kilometre by 1.2-kilometre island is a living
working community of more than 8,500 people
infinity pools and water villas of the luxury resort-only islands
it is the third most populous island in the Maldives
It is also one of the few Maldivian islands that is not surrounded by a coral lagoon
it is a volcanic bowl with the interior covered in tropical woodlands
This unique habitat within the Maldives is rich with endemic plant and animal species
It is a vital staging post for migratory birds – more than 167 bird species are found on the island
Maldivians consider Fuvahmulah the most beautiful of the 1,200 islands that make up the archipelago
It is a one-island atoll and the third-largest in size in the Maldives (excluding those such as the new Dhaalu international airport island which was extended by land reclaim)
Unlike most other islands it has working farms growing mangoes
pineapples and a range of vegetables such as aubergines and peppers
which separates Huvadhu Atoll from Addu Atoll
Fuvahmulah has long been a centre of tuna fishing
There is evidence that for more than 1,000 years
The rich waters surrounding the island are brimming with not just tuna but other pelagic species such as scalloped hammerhead sharks
melon-headed whales and a substantial population of tiger sharks
which extends for about three kilometres to the island’s southeast
These natural wonders led to UNESCO declaring Fuvamulah a biosphere reserve in 2020 – the only such reserve in the Maldives
Fishing is still the main occupation on the island
The locals use pole-and-line techniques which keep fish populations healthy and sustainable for future generations
which have decimated many other tuna fisheries
The islanders live by the philosophy of taking from the ocean and giving back to the ocean
This means that once they’ve gutted the fish
Herein lies the reason why this island is so special and could soon be joining the ranks of Cocos Island and the Galápagos Islands as one of the very best shark-diving destinations on the planet
a vast population of tiger sharks have become habituated to feeding on these scraps
A recent count identified more than 160 individuals
hanging around a ledge outside the harbour entrance
It is thought that hundreds more are regular visitors to the waters around the island
making it quite possibly the home of the largest concentration of tiger sharks on the planet
Add to that the bounty of the other pelagics found in the area
plus the remarkable reef-fish biodiversity
and you have a very special diving destination
His friends call him Lonu (the local word for saltwater) as Nasheed Ahmed was always in the ocean surfing (one of the advantages of not being surrounded by a coral lagoon is that the waves break onto the beach)
he decided he wanted to see the ever-present tiger sharks on their own terms
and jumped into the water at the edge of the harbour
Milling around the sandy rubble outside of the harbour were dozens of tiger sharks
He soon noticed that a significant number were heavily pregnant females
They seemed mildly interested in him but far more engaged with feeding on the tuna scraps
he opened the first dive centre on the island
He also founded the Fuvahmulah Marine Foundation to help conserve the island’s biodiversity
and organise and regulate the shark diving
Very few tourists and even fewer divers were visiting the island back then
like a saltwater version of the Kevin Costner movie Field of Dreams
first among dive guides working on the resort islands and
intrepid independent travellers started to arrive
but more and more divers are now coming to share Lonu’s passion
I visited Fuvahmulah with my wife Louise and four other divers with Ocean Culture Life in February this year
Our trip was guided by underwater photographer Jono Allen
The first time you jump into the water outside the harbour
you are glad you are with Lonu and have absorbed his thorough briefing
As you settle on the sand between eight to ten metres depth (depending on the tide)
it takes some getting used to when you realise you are surrounded by maybe five or even ten tiger sharks
The most tiger sharks we encountered at one time was 16
Evolution hasn’t equipped humans with 360-degree vision
leaving you feeling quite vulnerable at times
I began to take in the scene unfolding in front of me
There was a distinct hierarchy among the sharks
they left the ledge and slipped gently into the darkness; the younger sharks would come in to fight over the scraps
These younger sharks darted around more quickly than the larger
meaning you had to be more vigilant when they were nearby
you realised the sharks had distinct personalities
Some were hyper-inquisitive and would turn abruptly towards you to see if you were hiding fish from them
They seemed like puppies looking for scraps from the dinner table
Other would sedately glide past with a wary eye on you as if to say ‘Stay there
this is my kingdom and I’m tolerating you’
They behaved much like a strange dog might do if it came across you on a beach – bouncing up
At no point did I or my fellow divers believe we were on the menu
But this seemed to be friendly curiosity; as Jono put it
While the shark dives were the trip’s high point
The reefs were some of the healthiest I have seen in the Maldives in decades
skirting around triggerfish guarding their eggs
Fuvahmulah is going to change over the next few years
I can only hope that the efforts of Lonu and people such as Jono safeguard the island’s future
and that the divers who come to see the tiger sharks and all the other marine delights treat this extraordinary island with the respect it deserves
TAXONOMY: The tiger shark is a type of requiem shark of the Carcharhinidae family and the only living member of the genus Galeocerdo
Like all of the 270 different species of Carcharhiniformes
five-gill slits and has nictitating membrane over its eyes
SIZE & WEIGHT: Adults commonly reach between 3.25m and 4.25m in length and weigh up to 635 kilograms
There are records of exceptionally large females reaching in excess of 5m in length
One pregnant female caught in Australia measured 5.5m and weighed 1,524kg
It is the fourth-largest extant shark and the second-largest predatory shark (the great white being the largest).DISTRIBUTION & HABITAT: Found around the world in tropical and temperate waters
it stays on or near the continental shelves and ranges from the surface to depths of 900m
It is often found in estuaries and near harbours
Another common haunt is around coral atolls
it is found from Japan down to New Zealand
it is found throughout the East African coast
in the Maldives and around the shores of India and Southeast Asia
Tiger sharks can be seen in the Gulf of Mexico and in most of the Caribbean
and there are even recorded sightings in the Mediterranean
DIET: Larger tiger sharks tend to swim close inshore at night to feed
and head out to deeper waters during the day
such as around Fuvahmulah harbour in the Maldives
will attract large specimens during the day
They will eat sharks and rays and even other tiger sharks
A study in Australia found the remains of dugongs in 15 of 85 tiger shark stomachs examined
Another study in Hawaii found remains of terrestrial mammals
REPRODUCTION: Tiger sharks bear very large litters
with an average of between 26 to 33 pups born in spring to early summer
Mating occurs in spring – sometimes before pregnant females have given birth
They are the only species in its family that is ovoviviparous – that is
Sexual maturity is when the males reach 2.8m to 2.9m in length
GROWTH: Tiger sharks are fast-growing and reach maturity between four and 13 years
It is thought they live to at least 22 years
with some studies suggesting they may live as long as 37 years
RANGE: They are strong swimmers and have been known to undertake lengthy solo migrations
possibly for reasons of mating and pupping
Some tiger sharks have been tagged and discovered to journey thousands of kilometres a year
There are also semi-resident populations which barely move 100 kilometres each year
Most tiger sharks appear to be fairly solitary when mature
but there are aggregations when adults gather to feast on seasonal food supplies
such as the fledging of albatross chicks in the Pacific
THREAT: IUCN Red List – Near Threatened: In the Indian Ocean
tiger sharks have been intensely targeted by fishing
Its large fins are also prized in the Hong Kong markets
but there is some evidence that juvenile survival rates increase when adults are depleted.DIVING WITH TIGER SHARKS: Usually tiger sharks are unaggressive when encountered underwater by divers
and are commonly seen in shark-feed dives in The Bahamas and now Fuvahmulah
The island was only opened to tourists in 2016
it has had relatively rapid growth in tourism infrastructure
with small guesthouses and B&Bs opening up
a handful of dive centres and a smattering of restaurants and cafés catering for travellers
it is important to realise that this is not the luxury end of the travel market that anyone who has been to the Maldives in recent years will have experienced
A round-trip fare is approximately US$480 per person
The adventurous could take a cargo boat from Malé to Fuvahmulah
You can also reach Fuvahmulah by ferry from Addu Atoll in the south
Ferries are operated by Southern Transport Link and take between 45 and 90 minutes
Tickets need to be pre-booked and cost around US$20 each way
Addu Atoll is served by Gan International Airport in Addu City
plenty of Indian and Asian cuisine and even a touch of Mexico
New cafés and restaurants are opening up every month
so ask around about the latest places.Pebbles: Tables under thatched gazebos close to Thoondu Beach
serving traditional Maldivian delicacies along with Thai
Indian and other international cuisines.Royal Restaurant: Near the harbour
with a wide range of curries and seafood available
One of the best places to try Kattelhi fish stew – a local delicacy made from a rare deepwater fish that in the Maldives is only found on this island
Pre-order for the best fish.Real Breeze: Great Indian curries and the fish buns have a dedicated following.Runway 29: Small
Good ‘Hedhikaa’ menu of snacks and small meals
GETTING ABOUTAn asphalt road runs from the airport
much of the streets are unpaved and uneven
Most of the island is accessible via bicycles
available for hire at shops and some hotels and guesthouses
Motorbikes are also a popular method of transport and readily available for hire
There is also a very efficient taxi network with pre-booking only.
DON’T MISSThoondu Beach: Located towards the north of the island
It also has the only surf break in the Maldives
Surfing has recently become popular with the islanders and while there are as yet no schools
most hotels can arrange surf sessions with skilled locals
The Lakes: Bandaara Kilhi and Dhadimagi Kilhi form the island’s Nature Park
surrounded by stunning vegetation and endemic wildlife
MONEYThere are a few ATMs and a couple of banks which do foreign exchange
US dollars are widely accepted but in grocery stores
Some hotels and restaurants take credit cards
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The Indian Ocean archipelago has the third largest population in the world
1,000 individual oceanic manta rays have now been identified in Maldivian waters. According to the Maldives Manta Conservation Programme (MMCP) – the Manta Trust’s flagship project
which confirmed the 1,000th individual – this makes the Maldives home to the world’s third largest population of the species
The manta has been named ‘Anbu’ which means mango in Dhivehi
Oceanic mantas (Mobula birostris) are the world’s largest species of ray
Their wingspan can reach up to 7 metres – almost as long as an iconic London Routemaster bus – hence why they’re also known as giant mantas
can be identified by the markings on their underside
“Manta rays can be identified by the unique spot pattern on their belly, much like a human fingerprint,” explains Fauz Fath-hee, RahVeshi Programme Coordinator at MMCP
to take a photo of the spots on the manta rays’ bellies and we log each sighting in our ever-growing database.”
The charity has been studying mantas in the Maldives since 2005 but has ID photos going as far back as 1996
Citizen scientists and local scuba diving centres have also contributed photos in support of this important research
Although the Maldives is made up of 1,200 islands
nearly all sightings (85%) were from Fuvahmulah Atoll
a pelagic island in the south of the country known for its tiger shark dives.
With 1,000 individuals in its waters, the Maldives is home to the world’s third largest population of giant mantas (it also has the most reef mantas (Mobula alfredi) in the world, with over 5,000 individuals)
Ecuador has the largest oceanic manta population, “with 2,803 confirmed individuals but an estimated 22,316 individuals based on Proyecto Mantas Ecuador’s data,” says Fath-hee
The Pacific Manta Research Group has identified nearly 1,400 individuals in the Revillagigedo Archipelago in Mexico’s Eastern Pacific waters “so
the Maldives population is very close to becoming the second largest,” he adds
with the Indonesian archipelago of Raja Ampat in fourth with 588 individuals.
The researchers don’t yet fully understand why there are so many manta rays in the Maldives
“Unlike other oceanic manta ray aggregation sites
cleaning or courtship behaviour observed,” says Fath-hee who believes the rays may be transiting through because they are usually seen for only a day or two and re-sighting rates are low. “Where they are coming from and where they are going is still a mystery.” he says.
According to the IUCN Red List of Threatened Species, oceanic mantas are endangered so confirming there are at least 1,000 individuals gives conservationists hope
they are concerned the animals might be swimming on to Sri Lanka
“Oceanic manta rays are known to swim great distances
with some individuals traveling thousands of kilometres in search of food
has the world’s largest oceanic manta ray fishery and is believed to catch around 1,000 individuals each year
“There’s a high possibility that these rays could be moving between the protected waters of the Maldives and Sri Lanka’s waters where they face a high risk of being fished,” he says.
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